• DOI 10.31509/2658-607x-2020-3-4-1-90
  • УДК 574/577

Biodiversity and climate regulating functions of forests: current issues and prospects for research

N.V. Lukina1*, A.P. Geraskina1, A.V. Gornov1, N.E. Shevchenko1, A.V. Kuprin1,2, T.I. Chernov3, S.I. Chumachenko1,4, V.N. Shanin1,5,6, A.I. Kuznetsova1, D.N. Tebenkova1, M.V. Gornova1
1Center for Forest Ecology and Productivity of the Russian Academy of Sciences

117997 Moscow, Russian Federation, Profsoyuznaya st. 84/32 bldg. 14

2Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch of the Russian Academy of Sciences

690022 Vladivostok, Russian Federation, prospect 100 years of Vladivostok, 159

3Dokuchaev Soil Science Institute

397463 Moscow, Russian Federation, Pyzhyovskiy per., 7/2

4MB of Bauman Moscow State Technical University

141005 Mytischi, Moscow region, Russian Federation, 1st Institutskaya street, 1

5Institute of Physicochemical and Biological Problems in Soil Sciences of the Russian Academy of Science

142290 Puschino, Russian Federation, Institutskaya st., 2.

6Institute of Mathematical Problems of Biology RAS – the Branch of Keldysh Institute of Applied Mathematics of the Russian Academy of Sciences

142290 Puschino, Russian Federation, Prof. Vitkevich st., 1 


*E-mail: nvl07@yandex.ru

Received 30.11.2020

Accepted 22.12.2020

The problem of assessing the impact of biodiversity on the climate-regulating functions of forests has fundamental character and great importance for sustainable forest management in the context of global climate change. On the one hand, climate changes affect biodiversity, on the other hand, it is biodiversity, as a provider of all ecosystem functions, underlies the mechanisms of adaptation to these changes. This paper aims to discuss scientific questions about the links between biodiversity and climate-regulating functions of forests, and to outline the prospects for these studies. It is shown that studies of the influence of plant and animal species – ecosystem engineers on forest ecosystem’s functioning, including climate-regulating processes and functions, are quite numerous. However, studies of the combined effects of the diversity of biota belonging to different trophic levels and groups on climate-regulating functions of forests of different types/different stages of succession are not carried out. In such studies, it is important to take into account both taxonomic, including genetic, and functional biodiversity as well as structural diversity of forests. Various concepts of forest management taking into account the conservation and restoration of biodiversity are considered. An important aspect of this problem is the assessment and prediction of relationships (synergy or trade-offs) between climate-regulating and other ecosystem functions of forests with different levels of biodiversity functioning in natural conditions and under the combined impact of natural and anthropogenic factors, including climate change, fires, and forestry regimes. It is shown that a promising approach to assessing and predicting the dynamics of relationships between different ecosystem functions of forests is the integration of mathematical models.

Key words: forest ecosystems, taxonomic biodiversity, functional biodiversity, structural biodiversity, ecosystem functions, adaptation to climate change


Abaturov B.D., Mlekopitayushchie kak komponent ekosistem (na primere rastitel’noyadnyh mlekopitayushchih v polupustyne) (Mammals as a component of ecosystems (on the example of herbivorous mammals in a semi-desert), Moscow: Nauka, 1984, 288 p.

Abaturov B.D., Smirnov K.A., Formirovanie drevostoev na vyrubkah v lesah s raznoj plotnost’yu populyacii losya (Formation of stands in clearings in forests with different densities of elk populations), Byulleten’ MOIP. Otdel biologicheskij, 1992, Vol. 97, No. 3, pp. 3-12.

Aerts R., Nitrogen partitioning between resorption and decomposition pathways: a trade-off between nitrogen use efficiency and litter decomposibility? Oikos, 1997, Vol. 80, No 3, pp. 603-606.  

Alban D.H., Berry E.C., Effects of earthworm invasion on morphology, carbon, and nitrogen of a forest soil, Appl. Soil Ecol., 1994, Vol. 1, pp. 243-249.

Alejnikov A.A., Sostoyanie populyacii i sredopreobrazuyushchaya deyatel’nost’ bobra evropejskogo na territorii zapovednika «Bryanskij les» i ego ohrannoj zony (The state of the population and the environment-transforming activity of the European beaver on the territory of the reserve “Bryansk Les” and its buffer zone, Candidate’s biol. sci. thesis), Tol’yatti: Institut ekologii Volzhskogo bassejna RAN, 2010, 22 p.

Alekseev E.V., Tipy ukrainskogo lesa. Pravoberezh’e (Types of Ukrainian forest. Right bank), Kiev, 1925, 64 p.

Anan’eva N.D., Polyanskaya L.M., Stol’nikova E.V., Zvyagincev D.G., Sootnoshenie biomassy gribov i bakterij v profile lesnyh pochv (Fungal to bacterial biomass ratio in the forest soil profile), Izvestiya Rossijskoj akademii nauk. Seriya biologicheskaya, 2010, No. 3, pp. 308-317.

Anderegg W., Konings A., Trugman A.Yu.K., Bowling D., Karp D., Pacala S., Sperry J., Sulman B., Can plant functional diversity buffer forest ecosystem responses to drought? Geophysical Research Abstracts, 2018, Vol. 20, p. 8935.

Ansink E., Hein L., Hasund K.P., To value functions or services? An analysis of ecosystem valuation approaches, Environmental Values, 2008, Vol. 17, No 4, pp. 489-503.

Antonec N.V., Osobennosti royushchej deyatel’nosti dikogo kabana v poemnyh dubravah lesostepnoj i stepnoj zon (Features of burrowing activity of wild boar in the floodplain oak forests of the forest-steppe and steppe zones), Zapovidna sprava v Ukraїni, 1998, Vol. 4, Issue 2, pp. 18-24.

Artemkina N.A., Orlova, M.A., Lukina N.V. Micromosaic Structure of Vegetation and Variability of the Chemical Composition of L Layer of the Litter in Dwarf Shrub–Green Moss Spruce Forests of the Northern Taiga, Contemporary Problems of Ecology, 2018, Vol. 11, No 7, pp. 754-761.

Ashwood F., Vanguelova E.I., Benham S., Butt K.R., Developing a systematic sampling method for earthworms in and around deadwood, Forest Ecosystems, 2019, Vol. 6, No 33, pp. 1-12.

Augusto L., Ranger J., Binkley D., Rothe A., Impact of several common tree species of European temperate forests on soil fertility, Annals of Forest Science, 2002, No 59, pp. 233-253

Baeten L., Bruelheide H., van der Plas F., Kambach S., Ratcliffe S. …, & Scherer‐Lorenzen M., Identifying the tree species compositions that maximize ecosystem functioning in European forests, Journal of Applied Ecology, 2019,  Vol. 56, No 3, pp. 733-744.

Bagyaraj D.J., Ashwin R., Soil biodiversity: role in sustainable horticulture, Biodivers. Hortic. Crops., 2017, Vol. 5, pp. 1-18.

Bahmet O.N., Zapasy ugleroda v pochvah sosnovyh i elovyh lesov Karelii (Carbon storages in soils of pine and spruce forests in Karelia), Lesovedenie, 2018, No. 1, pp. 48-55.

Balodis M.M., Lesoekologicheskie aspekty bobrovogo hozyajstva v antropogennom landshafte (Forest-ecological aspects of beaver farming in the anthropogenic landscape), Lesovedenie, 1990, No. 1, pp. 29-37.

Bang H.S., Lee J.-H., Kwon O.S., Na Y.E., Jang Y.S., Kim W.H. Effects of paracoprid dung beetles (Coleoptera: Scarabaeidae) on the growth of pasture herbage and on the underlying soil, Applied soil ecology, 2005, Vol. 29, pp. 165-171.

Barber N.A., Hosler S.C., Whiston P., Jones H.P., Initial Responses of Dung Beetle Communities to Bison Reintroduction in Restored and Remnant Tallgrass, Natural Areas Journal, 2019, Vol. 39, No 4, pp. 420-428.

Bauhus J., Baber K., Müller J., Dead Wood in Forest Ecosystems, Oxford: Oxford University Press, 2018, pp. 1-16.

Baum C., El-Tohamy W., Gruda N., Increasing the productivity and product quality of vegetable crops using arbuscular mycorrhizal fungi: a review, Scientia Horticulturae, 2015, Vol. 187, pp. 131-141.

Begon M., Harper J.L., Townsend C.R., Ecology: Individuals, Populations and Communities, Oxford: Blackwell Scientific Publications, 1986, 1068 p.

Bender S.F., Wagg C., van der Heijden M.G., An underground revolution: biodiversity and soil ecological engineering for agricultural sustainability, Trends in ecology & evolution, 2016, Vol. 31, No. 6, pp. 440-452.

Berg B., Litter decomposition and organic matter turnover in northern forest soils, Forest Ecology and Management, 2000, Vol. 133, pp. 13-22.

Beringer J., Hutley L.B., Hacker J.M., Neininger B. Patterns and processes of carbon, water and energy cycles across northern Australian landscapes: from point to region, Agricultural and Forest Meteorology, 2011, Vol. 151, No 11, pp. 1409-1416.

Binkley D., Giardina Ch., Why do trees affect soils? The Warp and Woof of tree-soil interactions, Biogeochemistry, 1998, No 42, pp. 89-106.

Bityuckij N.P., Solov’eva A.N., Lukina E.I., Olejnik A.S., Zavgorodnyaya Yu.A., Demin V.V., Byzov B.A., Ekskrety dozhdevyh chervej – stimulyator mineralizacii soedinenij azota v pochve (Stimulating effect of earthworm excreta on the mineralization of nitrogen compounds in soil), Pochvovedenie, 2007, No. 4, pp. 468-473.

Bobek B., Perzanowski K., Energy and matter flow through ungulates [in:] Forest Ecosystems in Industrial Regions, W. Grodziriski, J. Weiner, P.F. Maycock (Eds.), Berlin: Springer-Verlag, 1984, pp. 121-125.

Bobkova K.S., Osipov A.F., Krugovorot ugleroda v sisteme “fitocenoz-pochva” v chernichno-sfagnovyh sosnyakah srednej tajgi Respubliki Komi (Carbon cycling in system “phytocenosis-soil” in bilberry-sphagnum pine forests of the middle taiga (Republic of Komi), Lesovedenie, 2012, No. 2, pp. 11-18.

Bobrovskij M.V., Lesnye pochvy Evropejskoj Rossii: bioticheskie i antropogennye faktory formirovaniya (Forest soils in European Russia: biotic and anthropogenic factors in pedogenesis), Moscow: Tovarishhestvo nauchnyh izdanij KMK, 2010, 359 p.

Bognounou F., Hulme P.E., Oksanen L., Suominen O., Olofsson J., Role of climate and herbivory on native and alien conifer seedling recruitment at and above the Fennoscandian tree line, Journal of Vegetation Science, 2018, Vol. 29, No 4, pp. 573-584.

Bohlen P.J., Pelletier D.M., Groffman P.M., Fahey T.J., Fisk M.C., Influence of earthworm invasion on redistributionand retention of soil carbon and nitrogen in northern temperate forests, Ecosystems, 2004, Vol. 7, pp. 13-27.

Bolton B., Identification guide to the ant genera of the world, Harvard University Press, 1994, 222 p.

Borges J.G., Nordström E.M., Garcia-Gonzalo J., Hujala T., Trasobares A. (Eds.), Computer-based tools for supporting forest management. The experience and the expertise world-wide, Report of Cost Action FP 0804 Forest Management Decision Support Systems (FORSYS), Sveriges Lantbruks universitet. Institutionen för skoglig resurshushållning, Umeå, 2014.

Bottinelli N., Capowiez Y., Earthworm ecological categories are not functional groups, Biology and Fertility of Soils, 2020, pp. 1-3.

Bouche M.B., Lombriciens de France: écologie et systématique, Paris: Institut national de la recherche agronomique, 1972, 671 p.

Boze B.G., Hernandez A.D., Huffman M.A., Moore J., Parasites and dung beetles as ecosystem engineers in a forest ecosystem, Journal of insect behavior, 2012. Vol. 25, No 4, pp. 352-361.

Bradford M.A., Wood S.A., Bardgett R.D., Black H.I., Bonkowski M., Eggers T., Jones T.H., Discontinuity in the responses of ecosystem processes and multifunctionality to altered soil community composition, Proceedings of the National Academy of Sciences, 2014, Vol. 111, No 40, pp. 14478-14483.

Brown J., Scholtz C.H., Janeau J.-L., Grellier S., Podwojewski P., Dung beetles (Coleoptera: Scarabaeidae) can improve soil hydrological properties, Applied soil ecology, 2010, Vol. 46, pp. 9-16.

Brussaard L., Soil fauna, guilds, functional groups and ecosystem processes, Applied soil ecology, 1998, Vol. 9, No 1-3, pp. 123-135.

Bugmann H., A review of forest gap models, Climatic Change, 2001, Vol. 51, pp. 259-305.

Bulavincev V.I., Formirovanie naseleniya melkih pozvonochnyh na territoriyah, narushennyh otkrytymi razrabotkami poleznyh iskopaemyh (Formation of the population of small vertebrates in territories disturbed by opencast mining of minerals), Zoologicheskij zhurnal, 1979, Vol. 58, No. 3, pp. 1884-1887.

Burtelow A.E., Bohlen P.J., Groffman P.M., Influence of exotic earthworm invasion on soil organic matter, microbial biomass and denitrification potential in forest soils of the northeastern United States, Appl. Soil Ecol., 1998, Vol. 9, pp. 197-202.

Buse J., Ranius T., Assmann T., An endangered longhorn beetle associated with old oaks and its possible role as an ecosystem engineer, Conservation Biology, 2008a, Vol. 22, pp. 329-337.

Buse J., Zabransky P., Assmann T., The xylobiontic beetle fauna of old oaks colonised by the endangered longhorn beetle Cerambyx cerdo Linnaeus, 1758 (Coleoptera: Cerambycidae), Mitt. Dtsch. Ges. Allg. Angew. Entomol., 2008b, Vol. 16, pp. 109-112.

Butt K.R., Earthworms in soil restoration: lessons learned from United Kingdom case studies of land reclamation, Restoration Ecology, 2008, Vol. 16, No 4, pp. 637-641.

Byhovec S.S., Komarov A.S., Prostoj statisticheskij imitator klimata pochvy s mesyachnym shagom (Simple statistical simulator of soil climate with a monthly step), Pochvovedenie, 2002, No. 4, pp. 443-452.

Byrnes J., Lefcheck J.S., Gamfeldt L., Griffin J.N., Isbell F., Hector A., Multifunctionality does not imply that all functions are positively correlated, Proceedings of the National Academy of Sciences of the United States of America, 2014, Vol. 111, No 51, pp. e5490.

Byzov B.A., Zoomikrobnye vzaimodejstviya v pochve (Zoomicrobial interactions in soil), Moscow: GEOS, 2005, 213 p.

Cadotte M.W., Carscadden K., Mirotchnick N., Beyond species: functional diversity and the maintenance of ecological processes and services, Journal of Applied Ecology, 2011, Vol. 48, pp. 1079-1087.

Cajander A.K., The theory of forest types, Acta Philosophica Fennica, 1926, Vol. 29, pp. 1-108.

Cao J., Wang C., Dou Z., Liu M., Ji D., Hyphospheric impacts of earthworms and arbuscular mycorrhizal fungus on soil bacterial community to promote oxytetracycline degradation, Journal of hazardous materials, 2018, Vol. 341, pp. 346-354.

Cardinale B.J., Matulich K.L., Hooper D.U., Byrnes J.E., Duffy E., Gamfeldt L., … & Gonzalez A., The functional role of producer diversity in ecosystems, American journal of botany, 2011, Vol. 98, No 3, pp. 572-592.

Carlile M.J., Watkinson S.C., Gooday G.W., Fungal diversity, The fungi, Academic Press, 2001, 2nd ed., pp. 11-84.

Chaer G., Fernandes M., Myrold D., Bottomley P., Comparative resistance and resilience of soil microbial communities and enzyme activities in adjacent native forest and agricultural soils, Microbial ecology, 2009, Vol. 58, pp. 414-424.

Chambers J.Q., Negron-Juarez R.I., Marra D.M., Di Vittorio A., Tews J., Roberts D., … & Higuchi N., The steady-state mosaic of disturbance and succession across an old-growth Central Amazon forest landscape, Proceedings of the National Academy of Sciences, 2013. Vol. 110, No 10, pp. 3949-3954.

Chernov I.YU., Marfenina O.E., Adaptivnye strategii gribov v svyazi s osvoeniem nazemnyh mestoobitanij (Adaptive strategies of fungi in connection with the development of terrestrial habitats), In: Paleopochvy i indikatory kontinental’nogo vyvetrivaniya v istorii biosfery (Fossil soil and indicators of continental weathering in the history of the biosphere), Moscow: PIN RAN, 2010, pp. 95-111.

Chernova N.M., Ekologicheskie sukcessii pri razlozhenii rastitel’nyh ostatkov (Ecological successions during decomposition of plant residues), Moscow: Nauka, 1977, 199 p.

Chertov O.G., Komarov A.S., Nadporozhskaya M.A., Bykhovets S.S., Zudin S.L., ROMUL – a model of forest soil organic matter dynamics as a substantial tool for forest ecosystem modelling, Ecological Modelling, 2001, Vol. 138, pp. 289-308.

Chertov O., Komarov A., Shaw C., Bykhovets S., Frolov P., Shanin, V., … & Shashkov M., Romul_Hum–a model of soil organic matter formation coupling with soil biota activity. II. Parameterisation of the soil food web biota activity, Ecological Modelling, 2017a, Vol. 345, pp. 125-139.

Chertov O., Shaw C., Shashkov M., Komarov A., Bykhovets S., Shanin V., …  & Zubkova E., Romul_Hum model of soil organic matter formation coupled with soil biota activity. III. Parameterisation of earthworm activity, Ecological Modelling, 2017b, Vol. 345, pp. 140-149.

Cheynier V., Comte G., Davies K.M., Lattanzio V., Martens S., Plant phenolics: recent advances on their biosynthesis, genetics, and ecophysiology, Plant physiology and biochemistry, 2013, Vol. 72, pp. 1-20.

Chumachenko S.I., Korotkov V.N., Palenova M.M., Politov D.V., Simulation modelling of long-term stand dynamics at different scenarios of forest management for conifer – broad-leaved forests, Ecol. Modeling., 2003, Vol. 170, pp. 345-361.

Chumachenko S.I., Smirnova O.V., Modelirovanie sukcesionnoj dinamiki nasazhdenij (Modeling of succession dynamics of forest stands), Lesovedenie, 2009, No. 6, pp. 3-17.

Cornelissen J.H., Lang S.I., Soudzilovskaia N.A., During H.J., Comparative cryptogam ecology: a review of bryophyte and lichen traits that drive biogeochemistry, Annals of botany, 2007a, Vol. 99, No. 5, pp. 987-1001.

Cornelissen J.H.C., van Bodegom P.M., Aerts R., Callaghan T.V., van Logtestijn R.S.P., … & Team M.O.L., Global negative vegetation feedback to climate warming responses of leaf decomposition rates in cold biomes, Ecological Letters, 2007b, Vol. 10, pp. 619-627.

Coulis M., Fromin N., David J.F., Gavinet J., Clet A., Devidal S., … & Hättenschwiler S., Functional dissimilarity across trophic levels as a driver of soil processes in a Mediterranean decomposer system exposed to two moisture levels, Oikos, 2015, Vol. 12, No 10, pp. 1304-1316.

Cromsigt J.P., Kemp Y.J., Rodriguez E., Kivit H., Rewilding Europe’s large grazer community: how functionally diverse are the diets of European bison, cattle, and horses? Restoration Ecology, 2018, Vol. 26, No 5, pp. 891-899.

Crowther T.W., Todd-Brown K.E.O., Rowe C.W., Wieder W.R., Carey J.C., Machmuller M.B., … & Bradford M.A., Quantifying global soil carbon losses in response to warming, Nature. 2016, Vol. 540, pp.104-108.

Dauber J., Wolters V., Microbial activity and functional diversity in the mounds of three different ant species, Soil Biology & Biochemistry, 2000, Vol. 32, pp. 93-99.

De Groot R.S., Alkemade R., Braat L., Hein L., Willemen L., Challenges in integrating the concept of ecosystem services and values in landscape planning, management and decision making, Ecological complexity, 2010, Vol. 7, No 3, pp. 260-272.

Demakov Yu.P., Diagnostika ustojchivosti lesnyh ekosistem: metodologicheskie i metodicheskie aspekty (Diagnostics of sustainability of forest ecosystems: methodological and methodical aspects), Yoshkar-Ola, 2000, 416 p.

Dezhkin V.V., D’yakov Yu.V., Safonov V.G., Bobr (Beaver), Мoscow: Agropromizdat, 1986, 256 с.

Dinesman L.G., Vliyanie dikih mlekopitayushchih na formirovanie drevostoev (The influence of wild mammals on the formation of forest stands), Moscow: AN SSSR, 1961, 165 p.

Dixon R.A., Paiva N.L., Stress-lnduced Phenylpropanoid Metabolism, The Plant Cell, 1995, Vol. 7, pp. 1085-1097.

Dmitrienko V.K., Lyudvig N.L., Vliyanie severnogo lesnogo murav’ya (Formica aquilonia Yarrov.) na vidovoj sostav i razvitie rastenij vozle muravejnika (Influence of the northern forest ant (Formica aquilonia Yarrov.) on the species composition and development of plants near the anthill), Materialy 12 Vserossijskogo mirmekologicheskogo simpoziuma (Materials of the 12th All-Russian Mirmecological Symposium), Novosibirsk, 2005, pp. 219-222.

Donlan J.C., Berger J., Bock C.E., Bock J.H., Burney D.A., Estes J.A., … & Soule M.E., Pleistocene rewilding: an optimistic agenda for twenty-first century conservation, The American Naturalist, 2006, Vol. 168, No 5, pp. 660-681.

Drake H.L., Horn M.A., Earthworms as a transient heaven for terrestrial denitrifying microbes: a review, Engineering in Life Sciences, 2006, Vol. 6, No 3, pp. 261-265.

Dylis N.V., Struktura lesnogo biogeocenoza (The structure of the forest biogeocenosis) Komarovskie chteniya, Moscow: Nauka, 1969, Vol. 21, 28 p.

Dymina G.D., Luga Dal’nego Vostoka (Zejsko-Bureinskoe Priamur’e) (Meadows of the Far East (Zeisko-Bureinskoe Priamurye)), Novosibirsk: Nauka, 1985, 193 p.

Eisenhauer N., Schielzeth Н., Barnes A., Barry K., Bonn A., & Ferlian O., A multi- trophic perspective on biodiversity–ecosystem functioning research, Advances in Ecological Research, 2019, Vol. 61, pp. 1-54.

El-Komy M.H., Saleh A.A., Eranthodi A., Molan Y.Y., Characterization of novel Trichoderma asperellum isolates to select effective biocontrol agents against tomato Fusarium wilt, The Plant Pathology Journal, 2015, Vol. 31, pp. 50-60.

Ernst G., Henseler I., Felten D., Emmerling C., Decomposition and mineralization of energy crop residues governed by earthworms, Soil Biology and Biochemistry, 2009, Vol. 41, No 7, pp. 1548-1554.

EUFORGEN – European Forest Genetic Resources Programme, URL.: http://www.euforgen.org (December 14, 2020).

European Russian forests: Their current state and features of their history. Smirnova O.V., Bobrovsky M.V., Khanina L.G. (Eds.), Plant and Vegetation, Vol. 15, Springer, Dordrecht, 2017, 566 p.

Evstigneev O.I., Belyakov K.V., Vliyanie deyatel’nosti bobra na dinamiku rastitel’nosti malyh rek (na primere zapovednika “Bryanskij les”) (Influence of beaver activity on the dynamics of vegetation of small rivers (on the example of the “Bryansky forest”)) Byulleten’ MOIP. Otdel biologicheskij, 1997, Vol. 102, Issue 6, pp. 34-41.

Evstigneev O.I., Korotkov V.N., Braslavskaya T.Yu., Kaban i ciklicheskie mikrosukcessii v travyanom pokrove shirokolistvennyh lesov (Wild boar and cyclic micro-successions in the grass cover of broad-leaved forests), In: Biogeocenoticheskij pokrov Nerusso-Desnyanskogo poles’ya: mekhanizmy podderzhaniya biologicheskogo raznoobraziya (Biogeocenotic cover of the Nerusso-Desnyanskogo polesia: mechanisms for maintaining biological diversity), Bryansk: Zapovednik “Bryanskij les”, 1999, pp.131-142.

Evstigneev O.I., Rubashko G.E., Chernyj sadovyj muravej i ciklicheskie sukcessii v travyanom pokrove vnutrilesnyh polyan (Black garden ant and cyclic successions in the grass cover of in-forest glades), In: Biogeocenoticheskij pokrov Nerusso-Desnyanskogo poles’ya: mekhanizmy podderzhaniya biologicheskogo raznoobraziya (Biogeocenotic cover of the Nerusso-Desnyanskogo polesia: mechanisms for maintaining biological diversity), Bryansk: Zapovednik “Bryanskij les”, 1999, pp. 143-161.

Evstigneev O.I., Solonina O.V., Zubr i podderzhanie bioraznoobraziya lugov (na primere zapovednika Bryanskij les) (European bison and maintenance of biodiversity of meadows (on the example of nature reserve Bryansky forest)), Byulleten’ MOIP. Otdel biologicheskij, 2016, Vol. 121, Issue 2, pp. 59-65.

Evstigneev O.I., Voevodin Evstigneev O.I., Korotkov V.N., Murashev I.A., Voevodin P.V., Zoochory and peculiarities of forest community formation: a review, Russian Journal of Ecosystem Ecology, 2017, Vol. 2, No. 1, pp. 1-16.

Evstigneev O.I., Solonina O.V., Phytocoenotic portrait of the European Badger, Russian Journal of Ecosystem Ecology, 2020, Vol. 5, No 1, pp. 1-26.

P.V., Korotkov V.N., Murashev I.A., Zoohoriya i dal’nost’ raznosa semyan v hvojno-shirokolistvennyh lesah Vostochnoj Evropy (Zoochory and distance of seeds dissemination in coniferous-broad-leaved forests of Eastern Europe), Uspekhi sovremennoj biologii, 2013, Vol. 133, No. 4, pp. 392-400.

Ferlian O., Eisenhauer N., Aguirrebengoa M., Camara M., Ramirez‐Rojas I., Santos F., … & Thakur M.P., Invasive earthworms erode soil biodiversity: A meta‐analysis, Journal of Animal Ecology, 2018, Vol. 87, No 1, pp. 162-172.

Feuda R.,  Bannikova A.A., Zemlemerova E.D., Febbraro M.D., Loy A. …, & Colangelo P., Tracing the evolutionary history of the mole, Talpa europaea, through mitochondrial DNA phylogeography and species distribution modelling, Biological Journal of the Linnean Society, 2015, Vol. 114, No 3, pp. 495-512.

Feurdean A., Ruprecht E., Molnar Z., Hutchinsond S.M., Hickler T., Biodiversity-rich European grasslands: Ancient, forgotten ecosystems, Biological Conservation, 2018, Vol. 228, pp. 224-232.

Filser J., Faber J.H., Tiunov A.V., Brussaard L., Frouz J., De Deyn G., … & Jimenez J.J., Soil fauna: Key to new carbon models, Soil, 2016, Vol. 2, pp. 565-582.

Finer L., Jurgensen M.F., Domisch T., Kilpeläinen J., Neuvonen S., Punttila P., … & Niemelä P., The role of wood ants (Formica rufa group) in carbon and nutrient dynamics of a boreal Norway spruce forest ecosystem, Ecosystems, 2013, Vol. 16, No 2, pp. 196-208.

Formozov A.N., Zveri, pticy i ih vzaimosvyazi so sredoj obitaniya (Mammals and birds and their interrelations with the environment), Moscow: Nauka, 1976, 309 p.

Formozov A.N., Zveri, pticy i ih vzaimosvyazi so sredoj obitaniya (Mammals and birds and their interrelations with the environment), Moscow: LKI, 2010, 312 p.

Frąc M., Hannula S.E., Bełka M., Jędryczka M., Fungal biodiversity and their role in soil health, Frontiers in Microbiology, 2018, Vol. 9, p. 707.

Framstad E., Biodiversity, Carbon Storage and Dynamics of Old Northern Forests, Copenhagen: Nordic Council of Ministers, 2013, 130 p.

Frelich L.E., Blossey B., Cameron E.K., Davalos A., Eisenhauer N., Fahey T., … & Maerz J.C., Side‐swiped: ecological cascades emanating from earthworm invasions // Frontiers in Ecology and the Environment. 2019. Vol. 17. No. 9. P. 502-510.

Freschet G.T., Cornwell W.K., Wardle D.A., Elumeeva T.G., Liu W., Jackson B.G., Onipchenko V.G., Soudzilovskaia N.A., Tao J., Cornelissen J.H.C. Linking litter decomposition of above- and below-ground organs to plant–soil feedbacks worldwide, Journal of Ecology, 2013, Vol. 101, pp. 943-952.

Freschet G.T., Roumet C., Comas L.H., Weemstra M., Bengough A.G., Rewald B., … & Lukac M., Root traits as drivers of plant and ecosystem functioning: current understanding, pitfalls and future research needs, New Phytologist, 2020, in press.

Fridland V.M., Problemy geografii genezisa i klassifikacii pochv (Problems of the geography of genesis and classification of soils), Moscow: Nauka, 1986, 243 p.

Fridman V.S., Eremkin G.S., Zaharova N.Yu., Vozvratnaya urbanizaciya-poslednij shans na spasenie uyazvimyh vidov ptic Evropy? (Return urbanization – the last chance to endangered species of birds in Europe and others high-urbanised regions?), Russian Journal of Ecosystem Ecology, 2016, Vol. 1, No. 4, pp. 1-58.

Frouz J., Livečkova M., Albrechtova J., Chroňakova A., Cajthaml T., Pižl V., … & Šimačkova H., Is the effect of trees on soil properties mediated by soil fauna? A case study from post-mining sites, Forest Ecology and Management, 2013, Vol. 309, pp. 87-95.

Gadd G.M., Geomycology: biogeochemical transformations of rocks, minerals, metals and radionuclides by fungi, bioweathering and bioremediation, Mycological research, 2007, Vol. 111, No 1, pp. 3-49.

Galushin V.M., Kostin A.B., Kubareva N.YU., Mechnikova S.A., Romanov M.S., Znachenie mikrofragmentov lesnoj rastitel’nosti dlya sohraneniya raznoobraziya hishchnyh ptic v agrocenozah pravoberezh’ya verhnego Dona (Importance of micro-fragments of forest vegetation for preserving the diversity of birds of prey in agrocenoses of the right bank of the upper Don), Redkie vidy ptic Nechernozyomnogo centra Rossii (Rare bird species of the Non-Black Earth Center of Russia, Proc. Conf. Title), Moscow: MPGU, 1998, pp. 174-179.

Gamfeldt L., Hillebrand H., Jonsson P.R., Multiple functions increase the importance of biodiversity for overall ecosystem functioning, Ecology, 2008, Vol. 89, No 5, pp. 1223-1231.

Gamfeldt L., Snäll T., Bagchi R., Jonsson M., Gustafsson L., Kjellander P., … & Mikusiński G., Higher levels of multiple ecosystem services are found in forests with more tree species, Nature communications, 2013, Vol. 4, No 1, pp. 1-8.

Gardi C., Jeffery S., Soil Biodiversity, Brussels: European Commission, 2009, p. 27.

Gehr B., Hofer E.J., Ryser A., Vimercati E., Vogt K., Keller L.F., Evidence for nonconsumptive effects from a large predator in an ungulate prey? Behavioral Ecology, 2018, Vol. 29, No. 3, pp. 724-735.

Geraskina A.P., Problemy kolichestvennoj ocenki i ucheta faunisticheskogo raznoobraziya dozhdevyh chervej v lesnyh soobshchestvah (Problems of quantification and accounting faunal diversity of earthworms in forest communities), Russian Journal of Ecosystem Ecology, 2016, Vol. 2, No. 2, pp. 1-9.

Geraskina A.P., Kiseleva L.L., Karpachev A.P., Abadonova M.N., Vliyanie reintrodukcii zubrov na kompleksy dozhdevyh chervej nacional’nogo parka “Orlovskoe Poles’e” (Bison reintroduction influence on the earthworms complexes of the national park “Orlovskoey Polesye”), Russian Journal of Ecosystem Ecology, 2018, Vol. 3, No. 4, pp. 1-21.

Geraskina A.P., Restoration of earthworms community (Oligochaeta: Lumbricidae) at sand quarries (Smolensk oblast, Russia), Ecological Questions, 2019, Vol. 30, No 3, pp. 7-15.

Geraskina A.P., Smirnova O.V., Korotkov V.N., Kudrevatykh I.Yu., Productivity and content of macro- and microelements in the phytomass of ground vegetation of typical and unique taiga forests of the Northern Urals (example of spruce-fir forests of the Pechora Ilych Nature Reserve), Russian Journal of Ecosystem Ecology, 2020, Vol. 5, No 2, pp. 1-13.

Geras’kina A.P., Vliyanie dozhdevyh chervej raznyh morfo-ekologicheskih grupp na akkumulyaciyu ugleroda v lesnyh pochvah (Impact of earthworms of different morpho-ecological groups on carbon accumulation in forest soils), Voprosy lesnoj nauki, 2020, Vol. 3. No. 2, pp. 1-20.

Gilyarov M.S., Krivoluckij D.A., Zhizn’ v pochve (Life in soil), Мoscow: Molodaya gvardiya, 1985, 191 p.

Gilyarov M.S., Rol’ stepnyh gryzunov v proiskhozhdenii entomofauny i sornopolevoj rastitel’nosti (The role of steppe rodents in the origin of entomofauna and weed-field vegetation), Dokl. AN SSSR, 1951, Vol. 79, No. 4, pp. 69-71.

Gleixner G., Soil organic matter dynamics: a biological perspective derived from the use of compound-specific isotopes studies, Ecological Research, 2013, Vol. 28, No 5, pp. 683-695.

Golichenkov M.V., Novosyolov A.L., Marfenina O.E., Dobrovol’skaya T.G., Zakalyukina YU.V., Lapygina E.V., Zamolodchikov D.G., Mikrobiologicheskaya harakteristika muravejnikov Lasius niger (Microbiological characteristic of anthills of Lasius niger), Izvestiya Rossijskoj akademii nauk. Seriya biologicheskaya, 2011, No. 3, pp. 334-339.

Goncharov A.A., Struktura troficheskih nish v soobshchestvah pochvennyh bespozvonochnyh (mezofauna) lesnyh ekosistem, Dis. kand. biol. nauk (The structure of trophic niches in the communities of soil invertebrates (mesofauna) of forest ecosystems. Candidate’s thesis), Moscow: IPEE RAN im. Severcova A.N., 2014, 177 p.

Gornov A.V., Zoogennaya i fitogennaya mozaichnost’ i floristicheskoe raznoobrazie vlazhnyh lugov Nerusso-Desnyanskogo poles’ya (Zoogenic and phytogenic patterns and floristic diversity of wet meadows in the Nerusso-Desnyanskoe polesye), Byulleten’ MOIP. Otdel biologicheskij, 2011, Vol. 116, Issue 6, p. 64-69.

Gornov A.V., Rol’ royushchih zhivotnyh v podderzhanii floristicheskogo raznoobraziya lesnyh soobshchestv (The role of burrowing animals in maintaining the floristic diversity of forest communities), In: Raznoobrazie i dinamika lesnyh ekosistem Rossii (Diversity and dynamics of forest ecosystems in Russia), Vol. 2, Moscow: Tovarishhestvo nauchnyh izdanij KMK, 2013, pp. 265-276.

Gornov A.V., Rol’ kabanov v podderzhanii populyacij nekotoryh vidov lugovyh rastenij v Nerusso-Desnyanskom poles’e (The role of wild boars in reproduction populations of some species of meadow plants in the Nerusso-Desnyanskoye Polesye) Byulleten’ Bryanskogo otdeleniya Russkogo botanicheskogo obshchestva, 2014, No. 2 (4), pp. 25-30.

 Grafius D.R.,  Corstanje R.,  Siriwardena G.M.,  Plummer K.E.,   Harris J.A., A bird’s eye view: using circuit theory to study urban landscape connectivity for birds, Landscape Ecology,  2017, Vol. 32, pp. 1771-1787.

Greenfacts, URL: https://www.greenfacts.org/glossary/def/ecosystem-processes.htm (December 14, 2020).

Gusev A.A., Pitanie losya i izmenenie rastitel’nosti v lesostepnyh dubravah (Moose nutrition and changes in vegetation in forest-steppe oak forests), Byulleten’ MOIP. Otdel biologicheskij, 1983, Vol. 88, Issue 6, pp. 46-50.

Gusev A.A., Funkcional’naya rol’ dikih kopytnyh zhivotnyh v zapovednyh biogeocenozah (The functional role of wild hoofed mammals in reserved biogeocenoses), In: Rol’ krupnyh hishchnikov i kopytnyh v biocenozah zapovednikov (The role of large predators and hoofed mammals in biocenoses of reserves), Moscow, 1986, pp. 94-105.

Gustafsson L., Bauhus J., Asbeck T., Augustynczik A.L.D., Basile M., Frey J., … & Knuff A., Retention as an integrated biodiversity conservation approach for continuous-cover forestry in Europe, Ambio, 2020, Vol. 49, No 1, pp. 85-97.

Hacker A.L., Coblentz B.E., Habitat selection by mountain beavers recolonizing Oregon coast range clearcuts, The Journal of wildlife management, 1993, Vol. 57, No 4, pp. 847-853.

Hale C.M., Frelich L.E., Reich P.B., Changes in hardwood forest understory plant communities in response to European earthworm invasions, Ecology, 2006, Vol. 87, No 7, pp. 1637-1649.

Hanina L.G., Bobrovskij M.V., Smirnov V.E., Grozovskaya I.S., Romanov M.S., Lukina N.V., Isaeva L.G., Funkcional’nye gruppy vidov i mikrogruppirovki lesnogo napochvennogo pokrova dlya modelirovaniya ego dinamiki (Ground vegetation modeling through functional species groups and patches in the forest floor), Matematicheskaya biologiya i bioinformatika, 2015, No. 1, pp. 15-33.

Hannula S.E., Morrien E., de Hollander M., Shifts in rhizosphere fungal community during secondary succession following abandonment from agriculture, The ISME journal, 2017, Vol. 11, No 10, pp. 2294-2304.

Hansen B.B., Henriksen S., Aanes R., Sæther B.E., Ungulate impact on vegetation in a two-level trophic system, Polar Biology, 2007, Vol. 30, No 5, pp. 549-558.

Hansson K., Impact of tree species on carbon in forest soils, Doctoral Thesis, Swedish University of Agricultural Sciences, Uppsala, 2011, 56 p.

Hättenschwiler S., Tiunov A.V., Scheu S., Biodiversity and litter decomposition in terrestrial ecosystems, Annual Review of Ecology Evolution and Systematics, 2005, Vol. 36, pp. 191-218.

Hawksworth D.L., Lücking R., Fungal diversity revisited: 2.2 to 3.8 million species [in:] The fungal kingdom, J. Heitman, B.J. Howlett, P.W. Crous, E.H. Stukenbrock, T.Y. James, N.A.R. Gow (Eds.), Washington: ASM Press, 2017, pp. 79-95.

Hedwall P.O., Brunet J., Trait variations of ground flora species disentangle the effects of global change and altered land-use in Swedish forests during 20 years, Global change biology, 2016, Vol. 22, No 12, pp. 4038-4047.

Hobbie S.E., Effects of plant species on nutrient cycling, Trends in ecology & evolution, 1992, Vol. 7, pp. 336-339.

Hodasheva K.S., Eliseeva V.I., Rol’ pozvonochnyh zhivotnyh – potrebitelej vetoshnyh kormov v krugovorote zol’nyh elementov (na primere lesostepnyh dubrov) (The role of vertebrates – consumers of waste fodder in the cycle of ash elements (on the example of forest-steppe oak trees)), In: Sredoobrazuyushchaya deyatel’nost’ zhivotnyh (Environment-forming activity of animals), Moscow: Izd-vo MGU, 1970, pp. 52-54.

Högberg M.N., Högberg P., Extramatrical ectomycorrhizal mycelium contributes one‐third of microbial biomass and produces, together with associated roots, half the dissolved organic carbon in a forest soil, New Phytologist, 2002, Vol. 154, No 3, pp. 791-795.

Holbrook K.M., Home Range and Movement Patterns of Toucans: Implications for Seed Dispersal, Biotropica, 2011, Vol. 3, No 3, pp. 357-364.

Hölldobler B., The ant fauna of Finnish Lapland, Waldhygiene, 1960, Vol. 3, No 8, pp. 229-238.

Hölldobler B., Wilson E.O., The ants, 1990, Harvard University Press.

Holodova M.V., Belousova I.P., Potreblenie i usvoenie pitatel’nyh veshchestv i energii zubrami (Bison bonasus) (Consumption and assimilation of nutrients and energy by bison (Bison bonasus)), Zoologicheskij zhurnal, 1989, Vol. 68, Issue 12, pp. 107-117.

Hooper D.U., Chapin F.S., Ewel J.J., Hector A., Inchausti P., Lavorel S., Lawton J.H., Lodge D.M., Loreau M., Naeem S., Schmid B., Setälä H., Symstad A.J., Vandermeer J., Wardle D.A., Effects of biodiversity on ecosystem functioning: a consensus of current knowledge, Ecological Monographs, 2005, Vol. 75, No 1, pp. 3-35.

Huang W., Gonzalez G., Zou X., Earthworm abundance and functional group diversity regulate plant litter decay and soil organic carbon level: A global meta-analysis, Applied Soil Ecology, 2020, Vol. 150, pp. 1-15.

IPCC, Climate Change and Land: an IPCC special report on climate change, desertification, land degradation, sustainable land management, food security, and greenhouse gas fluxes in terrestrial ecosystems, P.R. Shukla, J. Skea, E. Calvo Buendia, V. Masson-Delmotte, H.-O. Pörtner…, & J. Malley (Eds.), 2019, 874 p.,  https://www.ipcc.ch/srccl/ (December 14, 2020).

Isaev A.C., Girs G.I., Vzaimodejstvie dereva i nasekomyh-ksilofagov (Interaction between tree and xylophagous insects), Novosibirsk: Nauka, 1975, 347 p.

Isaev A.C., Kiselev V.V., Vetrova V.N., Vliyanie massovogo razmnozheniya bol’shogo chernogo hvojnogo usacha na sostoyanie lesnyh biogeocenozov (Influence of mass reproduction of the great black coniferous barbel on the state of forest biogeocenoses), Problemy ekologicheskogo monitoringa i modelirovaniya ekosistem, Leningrad: Gidrometeoizdat, 1981, Vol. 4, pp. 20-31.

Isaev A.S., Sukhovolsky V.G., Tarasova O.V., Palnikova E.N., Kovalev A.V., Forest insect population dynamics, outbreaks and global warming effects, John Wiley & Sons, 2017, p. 304.

Jacob M., Viedenz K., Polle A., Thomas F.M., Leaf litter decomposition in temperate deciduous forest stands with a decreasing fraction of beech (Fagus sylvatica), Oecologia, 2010, Vol. 164, No 4, pp. 1083-1094.

Jacobsen R.M., Burner R.C., Olsen S.L., Skarpaas O., Sverdrup-Thygeson A., Near-natural forests harbor richer saproxylic beetle communities than those in intensively managed forests, Forest Ecology and Management, 2020, Vol. 466, pp. 118-124.

Jastrow J.D., Amonette J.E., Bailey V.L., Mechanisms controlling soil carbon turnover and their potential application for enhancing carbon sequestration, Climatic Change, 2007, Vol. 80. No 1-2, pp. 5-23.

Jayne B., Quigley M., Influence of arbuscular mycorrhiza on growth and reproductive response of plants under water deficit: a meta-analysis, Mycorrhiza, 2014, Vol. 24, pp. 109-119.

Jenny H., Role of the plant factor in the pedogenic functions, Ecology, 1958, Vol. 39, No 1, pp. 5-16.

Jones C.G., Lawton J.H., Shachak M., Organisms as ecosystem engineers [in:] Ecosystem management, F.B. Samson, F.L. Knopf (Eds.), New York: Springer, 1994, pp. 130-147.

Jones C.G., Lawton J.H., Shachak M., Positive and negative effects of organisms as physical ecosystem engineers, Ecology, 1997, Vol. 78, No 7, pp. 1946-1957.

Jurgensen M.F., Finer L., Domisch T., Kilpeläinen J., Punttila P., Ohashi M., … & Risch A.C., Organic mound‐building ants: their impact on soil properties in temperate and boreal forests, Journal of Applied Entomology, 2008, Vol. 132, No 4, pp. 266-275.

Kalyakin V.N., Turubanova S.A., Izmenenie vidovogo sostava i rasprostraneniya klyuchevyh vidov (edifikatorov) mamontovogo kompleksa Vostochnoj Evropy s pozdnego plejstocena do pozdnego golocena (Changes in the species composition and distribution of key species (edificators) of the mammoth complex of Eastern Europe from the late pleistocene to the late holocene), In: Vostochnoevropejskie lesa: istoriya v golocene i sovremennost’ (Eastern European forests: history in Holocene and contemporaneity), Moscow: Nauka, 2004, Vol. 1, pp. 96-117.

Karpachevskij L.O., Pestrota pochvennogo pokrova v lesnom biogeocenoze (Heterogeneity of soil cover in forest biogeocenosis), Moscow: Izd-vo Mosk. un-ta, 1977, 312 p.

Karpachevskij M., Aksenov D., Esipova E., Vladimirova N., Danilova I., Kobyakov K., Zhuravleva I., Malonarushennye lesnye territorii Rossii: sovremennoe sostoyanie i utraty za poslednie 13 let (Intact forest areas of Russia: current state and losses over the past 13 years) Ustojchivoe lesopol’zovanie, 2015, Vol. 42, No. 2, pp. 2-7.

Karpechko Yu.V., Gidrologicheskaya ocenka antropogennogo vozdejstviya na vodosbory v taezhnoj zone Evropejskogo Severa Rossii, Dis. dokt. geogr. nauk (Hydrological assessment of anthropogenic impact on catchments in the taiga zone of the European North of Russia. Doktor’s thesis), Saint Petersburg, 2004, 303 p.

Karpechko Yu.V., Kondrat’ev S.A., Rodionov V.Z., SHmakova M.V., Osobennosti formirovaniya ispareniya v razlichnyh po vozrastu, usloviyam proizrastaniya i produktivnosti lesah (Evaporation patterns in forests of different ages, site conditions and productivity levels), Gidrometeorologiya i ekologiya, 2020, No. 58, pp. 49-67.

Kazimirov N.I., Morozova R.M., Biologicheskij krugovorot veshchestv v el’nikah Karelii (Biological circulation of substances in the spruce forests of Karelia), Leningrad: Nauka, 1973, 175 p.

Kazimirov N.I., Volkov A.D., Zyabchenko S.S., Ivanchikov A.A., Morozova R.M., Obmen veshchestv i energii v sosnovyh lesah Evropejskogo Severa (Exchange of matters and energy in pine forests of the European North), Leningrad: Nauka, 1977, 304 p.

Kaz’min V.D., Smirnov K.A., Zimnee pitanie, kormovye resursy i troficheskoe vozdejstvie zubrov na lesnye fitocenozy Central’nogo Kavkaza (Winter food, forage resources and trophic impact of bison on forest phytocenoses of the Central Caucasus), Byulleten’ MOIP. Otdel biologicheskij, 1992, Vol. 97, No. 2, pp. 26-35.

Khanina L.G., Bobrovsky M.V., Komarov A.S., Mikhajlov A.V., Modeling dynamics of forest ground vegetation diversity under different forest management regimes, Forest Ecology & Management, 2007, Vol. 248, pp. 80-94.

Khlifa R., Paquette A., Messier C., Reich P.B., Munson A.D., Do temperate tree species diversity and identity influence soil microbial community function and composition?  Ecology and evolution, 2017, Vol. 7, No 19, pp. 7965-7974. 

Khlifa R., Angers D.A., Munson A.D., Understory Species Identity Rather than Species Richness Influences Fine Root Decomposition in a Temperate Plantation, Forests, 2020, Vol. 11, No 10, pp. 1091.  

Komarov A.S., Chertov O.G., Zudin S.L., Nadporozhskaya M.A., Mikhailov A.V., Bykhovets S.S., Zudina E.V., Zoubkova E.V., EFIMOD 2 – A model of growth and elements cycling of boreal forest ecosystems, Ecological Modelling, 2003, Vol. 170, pp. 373-392.

Komarov A., Chertov O., Bykhovets S., Shaw C., Nadporozhskaya M., Frolov P., … & Zubkova E., Romul_Hum model of soil organic matter formation coupled with soil biota activity. I. Problem formulation, model description, and testing, Ecological Modelling, 2017, Vol. 345, pp. 113-124. 

Kondrat’ev S.A., Karpechko Yu.V., SHmakova M.V., Vliyanie vyrubok lesa na stok i vynos biogennyh elementov s lesnyh vodosborov Karelii (po dannym matematicheskogo modelirovaniya) (Impact of forest cutting down on runoff and nutrient removal from forest catchments of Karelia (according to mathematical modeling)), Gidrometeorologiya i ekologiya, 2020, No. 59, pp. 51-66.

Korochkina L.N., Drevesnaya rastitel’nost’ v pitanii zubrov Belovezhskoj Pushchi (Woody vegetation in the diet of bison of Belovezhskaya Pushcha), Belovezhskaya Pushcha. Minsk, 1969a, pp. 120-126.

Korochkina L.N., Rajon obitaniya i stacial’noe razmeshchenie zubrov v Belovezhskoj pushche (Habitat and stationary distribution of bison in Belovezhskaya Pushcha), Belovezhskaya pushcha. Issledovaniya, Minsk: Uradzhaj, 1973, Issue 7, pp. 148-165.

Korochkina L.N., Vidovoj sostav lesnoj travyanistoj rastitel’nosti v pitanii zubrov Belovezhskoj Pushchi (Species composition of forest herbaceous vegetation in the diet of bison in Belovezhskaya Pushcha), Belovezhskaya Pushcha, Minsk, 1969b, pp. 204-221.

Korotkov V.N., Osnovnye koncepcii i metody vosstanovleniya prirodnyh lesov Vostochnoj Evropy (Basic concepts and methods of restoration of natural forests in Eastern Europe), Russian Journal of Ecosystem Ecology, 2017, Vol. 2, No. 1, DOI:10.21685/2500-0578-2017-1-1.

Kovalev I.V., Kovaleva N.O., Pul ligninovyh fenolov v pochvah lesnyh ekosistem (Pool of lignin phenols in soils of forest ecosystems), Lesovedenie, 2016, No. 2, pp. 148-160.

Kovaleva N.O., Kovalev I.V., Ligninovye fenoly v pochvah kak biomarkery paleorastitel’nosti (Lignin phenols in soils as biomarkers of paleovegetation), Pochvovedenie, 2015, No. 9, pp. 1073-1086.

Kozlo P.G., Stavrovskaya L.A., Vliyanie royushchej deyatel’nosti kabana (Sus scrofa L.) na travyanuyu rastitel’nost’ (Influence of burrowing activity of wild boar (Sus scrofa L.) on herbaceous vegetation), Zapovedniki Belorussii, Minsk, 1974, Issue 3, pp. 91-99.

Kozlovskaya L.S., Belous A.P., Izmenenie organicheskoj chasti rastitel’nyh ostatkov pod vliyaniem oligohet (Change of organic matter of plants rests under the influence of Oligochaetes), In: Vzaimootnosheniya lesa i bolota (The relationship between forest and swamp), Moscow: Nauka, 1967, pp. 43-55.

Kraus D., Krumm F., (Eds.) Integrative approaches as an opportunity for the conservation of forest biodiversity, Germany: European Forest Institute, 2013, 284 p.

Kudeyarov V.N., Zavarzin G.A., Blagodatskij S.A., Borisov A.V., Voronin P.YU. … & Chertov O.G., Puly i potoki ugleroda v nazemnyh ekosistemah Rossii (Pools and fluxes of carbon in terrestrial ecosystems in Russia), Moscow: Nauka, 2007, 315 p.

Kuijper D.P.J., Sahlén E., Elmhagen B., Chamaillé-Jammes S., Sand H., Lone K., Cromsigt, J.P.G.M., Paws without claws? Ecological effects of large carnivores in anthropogenic landscapes. Proceedings, Biological Sciences, 2016, Vol. 283, Article: 1841.

Kurakov A.V., Kharin S.A., Byzov B.A., Changes in the composition and physiological and biochemical properties of fungi during passage through the digestive tract of earthworms, Biological Bulletin, 2016, Vol. 43, pp. 290-299.

Kurcheva G.F., Rol’ pochvennyh zhivotnyh v razlozhenii i gumifikacii rastitel’nyh ostatkov (The role of soil animals in the decomposition and humification of plant residues), Moscow: Nauka, 1971, 156 p.

Kurek P., Kapusta P., Holeksa J., Burrowing by badgers (Meles meles) and foxes (Vulpes vulpes) changes soil conditions and vegetation in a European temperate forest, Ecological Research, 2014, Vol. 29, No 1, pp. 1-11.

Kurek P., Topsoil mixing or fertilization? Forest flora changes in the vicinity of badgers’ (Meles meles L.) setts and latrines, Plant and Soil, 2019, Vol. 437, pp. 1-2, pp. 327-340.

Kurkin K.A., Sistemnye issledovaniya dinamiki lugov (System studies of meadow dynamics), Moscow: Nauka, 1976, 284 p.

Kurz W.A., Dymond C.C., White T.M., Stinson G., Shaw C.H., Rampley G.J., Smyth C., Simpson B.N., Neilson E.T., Trofymow J.A., Metsaranta J., Apps M.J., CBM-CFS3: A model of carbon-dynamics in forestry and land-use change implementing IPCC standards, Ecological Modelling, 2009, Vol. 220, pp. 480-504.

Kutovaya O.V., Vliyanie dozhdevyh chervej (Oligochaeta, Lumbricidae) na biotu i organicheskoe veshchestvo dernovo-podzolistyh pochv pri raznyh sistemah zemlepol’zovaniya, Avtoref. dis kand. s-h. nauk (Influence of earthworms (Oligochaeta, Lumbricidae) on biota and organic matter of sod-podzolic soils under different land use system, Extended abstract of Candidate’s thesis), Moscow, 2012, 27 p.

Kuuluvainen T., Gap disturbance, ground microtopography, and the regeneration dynamics of boreal coniferous forests in Finland: a review, Annales Zoologici Fennici, Finnish Zoological Publishing Board, formed by the Finnish Academy of Sciences, Societas Biologica Fennica Vanamo, Societas pro Fauna et Flora Fennica, and Societas Scientiarum Fennica, 1994, pp. 35-51.

 Lacey  E.A., James L.P.,  Cameron G.N., (Eds). Life Underground, The biology of subterrainean rodents, The University of Chicago Press Book, 2000, 479 p.

Lachat T., Wermelinger B., Gossner M., Bussler H., Isacsson G., Müller J., Saproxylic beetles as indicator species for dead-wood amount and temperature in European beech forests, Ecological Indicators, 2012, Vol. 23, pp. 323-331.

Laganiere J., Paré D., Bergeron Y., Chen H.Y., Brassard B.W., Cavard X., Stability of soil carbon stocks varies with forest composition in the Canadian boreal biome, Ecosystems, 2013, Vol. 16, pp. 852-865.

Laine K.J., Niemelä P., The influence of ants on the survival of mountain birches during an Oporinia autumnata (Lep. Geometridae) outbreak, Oecologia, 1980, Vol. 47, No 1, pp. 39-42.

Lang S.I., Cornelissen J.H.C., Klahn T., van Logtestijn R.S.P., Broekman R., Schweikert W., Aerts R., An experimental comparison of chemical traits and litter decomposition rates in a diverse range of subarctic bryophyte, lichen and vascular plant species, Journal of Ecology, 2009, Vol. 97, No 5, pp. 886-900.

Le Bayon R.C., Bullinger-Weber G., Schomburg A., Turberg P., Schlaepfer R., Guenat C., Earthworms as ecosystem engineers: a review [in:] Earthworms: Types, Roles and Research (Ed. C.G. Horton). New York: Nova Science Publishers, 2017, pp. 129-178.

Lee K.E., Earthworms. Their ecology and relationships with soils and land use, Academic press (Harcourt Brace Jovanovich, Publishers), 1985, pp. 211-221.

Lee S.-G., Kim C., Kuprin A.V., Kang J.-H., Lee B.-W., Oh S.H., Lim J., Survey research on the habitation and biological information of Callipogon relictus Semenov in Gwangneung forest, Korea and Ussurisky nature reserve, Russia (Coleoptera, Cerambycidae, Prioninae), ZooKeys, 2018, No 792, pp. 45-68.

Lee S.Y., Foster R.C., Soil fauna and soil structure, Australian Journal of Soil Research, 1991, Vol. 29, pp. 745-775.

Levina R.E., Sposoby rasprostraneniya plodov i semyan (Distribution of fruits and seeds), Moscow, 1957, 360 p.

Liang J., Crowther T.W., Picard N., Wiser S., Zhou M., Alberti G., … & De-Miguel S., Positive biodiversity-productivity relationship predominant in global forests, Science, 2016, Vol. 354, No 6309, pp. 1-15.

Logofet D.O., Evstigneev O.I., Alejnikov A.A., Morozova A.O., Sukcessiya, vyzvannaya zhiznedeyatel’nost’yu bobra (Castor fiber L.): I. Uroki kalibrovki prostoj markovskoj modeli (Succession caused by beaver (Castor fiber L.) life activity: I. What is learnt from the calibration of a simple Markov model), ZHurnal obshchej biologii, 2014, Vol. 75, No. 2, pp. 95-103.

Logofet D.O., Evstigneev O.I., Alejnikov A.A., Morozova A.O., Sukcessiya, vyzvannaya zhiznedeyatel’nost’yu bobra (Castor fiber L.): II. Utochnennaya markovskaya model’ (Succession caused by beaver (Castor fiber L.) life activity: II. A refined Markov model), Zhurnal obshchej biologii, 2015, Vol. 76, No. 2, pp. 126-145.

Lord C.M., Wirebach K.P., Tompkins J., Bradshaw-Wilson C., Shaffer C.L., Reintroduction of the European bison (Bison bonasus) in central-eastern Europe: a case study, International Journal of Geographical Information Science, 2020, Vol. 34, No 8, pp. 1628-1647.

Louman B., Cifuentes M., Chacón M., REDD+, RFM, Development, and Carbon Markets, Forests, 2011, Vol. 2, No 1, pp. 357-372.

Lubbers I.M., Brussaard L., Otten W., Van Groenigen J.W., Earthworm‐induced N mineralization in fertilized grassland increases both N2O emission and crop‐N uptake, European Journal of Soil Science, 2011, Vol. 62, No 1, pp. 152-161.

Lubbers I.M., Gonzalez E.L., Hummelink E.W.J., Van Groenigen J.W., Earthworms can increase nitrous oxide emissions from managed grassland: a field study, Agriculture, ecosystems & environment, 2013, Vol. 174, pp. 40-48.

Lubbers I.M., Pulleman M.M., Van Groenigen J.W., Can earthworms simultaneously enhance decomposition and stabilization of plant residue carbon? Soil Biology and Biochemistry, 2017, Vol. 105, pp. 12-24.

Lubbers I.M., Berg M.P., De Deyn G.B., van der Putten W.H., van Groenigen J.W., Soil fauna diversity increases CO2 but suppresses N2O emissions from soil, Global change biology, 2020, Vol. 26, No 3, pp. 1886-1898.

Lugovaya D.L., Smirnova O.V., Zaprudina M.V., Aleynikov A.A., Smirnov V.E., Micromosaic structure and phytomass of ground vegetation in main types of dark conifer forests in the pechora–ilych state nature reserve, Russian Journal of Ecology, 2013, Vol. 44, No 1, pp. 3-10.

Lukina N.V., Nikonov V.V., Biogeohimicheskie cikly v lesah Severa v usloviyah aerotekhnogennogo zagryazneniya (Biogeochemical cycles in the forests of the North under conditions of airborne industrial pollution), In 2 parts, Part 1, Apatity: Izd-vo Kol’skogo nauchnogo centra RAN, 1996, 213 p. Part 2, Apatity: Izd-vo Kol’skogo nauchnogo centra RAN, 1996, 192 p.

Lukina N.V., Nikonov V.V., Pitatel’nyj rezhim lesov severnoj tajgi: prirodnye i tekhnogennye aspekty (Nutrient regime of northern taiga forests: natural and technogenic aspects), Apatity: Izd-vo Kol’skogo nauchnogo centra RAN, 1998, 316 p.

Lukina N.V., Orlova M.A., Isaeva L.G., Plodorodie lesnyh pochv kak osnova vzaimosvyazi pochva-rastitel’nost’ (Forest soil fertility: the base of relationships between soil and vegetation), Lesovedenie, 2010, No. 5, pp. 45-56.

Lukina N.V., Orlova M.A., Steinnes E., Artemkina N.A., Gorbacheva T.T., Smirnov V.E., Belova E.A., Mass-loss rates from decomposition of plant residues in spruce forests near the northern tree line subject to strong air pollution, Environmental Science and Pollution Research, 2017, Vol. 24, No 24, pp. 19874-19887.

Lukina N.V., Tikhonova E.V., Orlova M.A., Bakhmet O.N., Kryshen A.M. …& Zukert N.V., Associations between forest vegetation and the fertility of soil organic horizons in northwestern Russia, Forest ecosystems, 2019, Vol. 6, No 1, p. 34.

Lukina N., Kuznetsova A., Tikhonova E., Smirnov V., Danilova M., Gornov A., Bakhmet O., Kryshen A., Tebenkova D., Shashkov M., Knyazeva S., Linking Forest Vegetation and Soil Carbon Stock in Northwestern Russia, Forests, 2020, Vol. 11, No 9, p. 979.

Maccarthy J.W. Gap dynamics of forest trees: A review with particular attention to boreal forests, Environmental Reviews, 2001, Vol. 9, No 1, pp. 1-59.

Maes S.L., Perring M.P., Depauw L., Bernhardt-Römermann M., Blondeel H., … & Verheyen K., Plant functional trait response to environmental drivers across European temperate forest understorey communities, Plant Biology, 2020, Vol. 22, No 3, pp. 410-424.

Maestre F.T., Quero J.L., Gotelli N.J., Escudero A., Ochoa V., Delgado-Baquerizo M., García-Palacios P., Plant species richness and ecosystem multifunctionality in global drylands, Science, 2012, Vol. 335, No 6065, pp. 214-218.

Makarov M.I., Buzin I.S., Tiunov A.V., Malysheva T.I., Kadulin M.S., Koroleva N.E., Nitrogen isotopes in soils and plants of tundra ecosystems in the Khibiny Mountains, Eurasian Soil Science, 2019, Vol. 52, No 10, pp. 1195-1206.

Makeschin F., Earthworms (Lumbricidae: Oligochaeta): Important promoters of soil development and soil fertility [in:] Fauna in soil ecosystems. Recycling processes, nutrient fluxes and agricultural production. (Ed. G. Benckiser),1997, pp. 173-223.

Makkonen M., Berg M.P., Handa I.T., Hättenschwiler S., van Ruijven J., van Bodegom P.M., Aerts R., Highly consistent effects of plant litter identity and functional traits on decomposition across a latitudinal gradient, Ecology Letters, 2012, Vol. 15, pp. 1033-1041.

Mamkin V., Kurbatova J., Avilov V., Ivanov D., Kuricheva O., Varlagin A., Yaseneva I., Olchev A. Energy and CO2 exchange in an undisturbed spruce forest and clear-cut in the Southern Taiga, Agricultural and Forest Meteorology, 2019, Vol. 265, pp. 252-268.

Manakov K.N., Nikonov V.V., Biologicheskij krugovorot mineral’nyh elementov i pochvoobrazovanie v el’nikah Krajnego Severa (Biological cycle of mineral elements and pedogenesis in the spruce forests of the Far North), Leningrad: Nauka, 1981, 196 p.

Manning P., Plas F., Soliveres S., Allan E., Maestre F.T., Mace G., Fischer M., Redefining ecosystem multifunctionality, Nature ecology & evolution, 2018, Vol. 3, p. 427.

McDaniel J.P., Stromberger M.E., Barbarick K.A., Cranshaw W., Survival of Aporrectodea caliginosa and its effects on nutrient availability in biosolids amended soil, Applied soil ecology, 2013, Vol. 71, pp. 1-6.

Metodicheskie rekomendacii po vosproizvodstvu raznovozrastnyh shirokolistvennyh lesov evropejskoj chasti SSSR (na osnove populyacionnogo analiza) (Methodological recommendations for the reproduction of broad-leaved forests of different ages in the European part of the USSR (based on population analysis), O.V. Smirnova, R.V. Popadyuk, A.A. Chistyakova et al. (Eds.), Мoscow: VASKHNIL, 1989, 19 p.

Migge-Kleian S., McLean M.A., Maerz J.C., Heneghan L., The influence of invasive earthworms on indigenous fauna in ecosystems previously uninhabited by earthworms, Biological Invasions, 2006, Vol. 8, No 6, pp. 1275-1285.

Millennium Ecosystem Assessment. Ecosystems and Human Wellbeing: Synthesis. Washington, DC: Island Press. 2005. (URL: http://www.millenniumassessment.org/en/Reports.aspx#) (13.12.2020).

Mirkin B.M., Naumova L.G., Metod klassifikacii rastitel’nosti po Braun-Blanke v Rossii (Braun-Blanquet method of vegetation classification in Russia), ZHurnal obshchej biologii, 2009, Vol. 70, No. 1, pp. 66-77.

Mlekopitayushchie v nazemnyh ekosistemah (Mammals in terrestrial ecosystems), Moscow: Nauka, 1985, 289 p.

Moradi J., Vicentini F., Šimačková H., Pižl V., Tajovský K., Stary J., Frouz J., An investigation into the long-term effect of soil transplant in bare spoil heaps on survival and migration of soil meso and macrofauna, Ecological Engineering, 2018, Vol. 110, pp. 158-164.

Mori A.S., Isbell F., Fujii S., Makoto K., Matsuoka S., Osono T., Low multifunctional redundancy of soil fungal diversity at multiple scales, Ecology letters, 2016, Vol. 19, pp. 249-259.

Mori A.S., Lertzman K.P., Gustafsson L., Biodiversity and ecosystem services in forest ecosystems: a research agenda for applied forest ecology, Journal of Applied Ecology, 2017, Vol. 54, No 1, pp. 12-27.

Mouillot D., Villéger S., Scherer-Lorenzen M., Mason N.W., Functional structure of biological communities predicts ecosystem multifunctionality, PloS one, 2011, Vol. 6, No 3, p. e17476.

Mueller K.E., Eissenstat D.M., Hobbie S.E., Oleksyn J., Jagodzinski A.M., Reich P.B., Chawick O.A., Chorover J., Tree species effects on coupled cycles of carbon, nitrogen and acidity in mineral soils at a common garden experiment, Biogeochemistry, 2015, Vol. 111, No 1-3, pp. 601-614.

Muller R.N., Nutrient relations of the herbaceous layer in deciduous forest ecosystems [in:] The Herbaceous Layer in Forests of Eastern North America, F.S. Gilliam, M.R. Roberts (Eds.), New York: Oxford University Press, 2003, pp. 15-37.

Muscolo A., Bagnato S., Sidari M., Mercurio R., A review of the roles of forest canopy gaps, Journal of Forestry Research, 2014, Vol. 25, No 4, pp. 725-736.

Naeem S., Loreau M., Inchausti P., Biodiversity and ecosystem functioning: the emergence of a synthetic ecological framework, Biodiversity and ecosystem functioning: synthesis and perspectives, 2002, pp. 3-11.

Nakonechnyj N.V., Ekologicheskoe znachenie hodov obyknovennogo krota (Talpa europaea L., 1758) v formirovanii faunisticheskih kompleksov v lesnoj zone Zapadnoj Sibiri, Dis. kand. biol. nauk (The ecological significance of the passages of the common mole (Talpa europaea L., 1758) in the formation of faunal complexes in the forest zone of Western Sibiria. Candidate’s thesis), Surgut: SurGU, 2013, 176 p.

Ndiade-Bourobou D., Hardy O.J., Favreau B., Moussavou H., Nzengue E., Mignot A., Bouvet J.M., Long-distance seed and pollen dispersal inferred from spatial genetic structure in the very low-density rainforest tree, Baillonella toxisperma Pierre, in Central Africa, Molecular Ecology, 2010, Vol. 19, No 22, pp. 4949-4962.

Nebert L.D., Bloem J., Lubbers I.M., van Groenigen J.W., Association of earthworm-denitrifier interactions with increased emission of nitrous oxide from soil mesocosms amended with crop residue, Applied and Environmental Microbiology, 2011, Vol. 77, No 12, pp. 4097-4104.

Nemcev A.S., Rautian G.S., Puzachenko A.YU., Sipko T.P., Kalabushkin B.A., Mironenko I.V., Zubr na Kavkaze (Bison in the Caucasus), Majkop: Kachestvo, 2003, 292 p.

Nichols E., Spector S., Louzada J., Larsen T., Amezquita S., Favila M.E., Network T.S.R., Ecological functions and ecosystem services provided by Scarabaeinae dung beetles, Biological conservation, 2008, Vol. 141, No 6, pp. 1461-1474.

Niklasson M., Granström A., Numbers and sizes of fires: long-term spatially explicit fire history in a Swedish boreal landscape, Ecology, 2000, Vol. 81, pp. 1484-1499.

Nikonov V.V., Lukina N.V., Biogeohimicheskie funkcii lesov na severnom predele rasprostraneniya (Biogeochemical functions of forests at the northern limit of distribution), Apatity: Izd-vo Kol’skogo nauchnogo centra RAN, 1994, 315 p.

Novara A., Rühl J., La Mantia, T., Gristina L., La Bella, S., Tuttolomondo T., Litter contribution to soil organic carbon in the processes of agriculture abandon, Solid Earth, 2015, Vol. 6, No 2, pp. 425-432.

Nummi P., Kattainen S., Ulander P., Hahtola A., Bats benefit from beavers: a facilitative link between aquatic and terrestrial food webs, Biodiversity and Conservation, 2011, Vol. 20, No 4, pp. 851-859.

Nummi P., Holopainen S., Restoring wetland biodiversity using research: Whole‐community facilitation by beaver as framework, Aquatic Conservation: Marine and Freshwater Ecosystems, 2020, Vol. 30, No 9, pp. 1798-1802.

Nygaard P.H., Strand L.T., Stuanes A.O., Gap formation and dynamics after long‐term steady state in an old‐growth Picea abies stand in Norway: Above‐and belowground interactions, Ecology and evolution, 2018, Vol. 8, No 1, pp. 462-476.

O’Connor M.I., Gonzalez A., Byrnes J.E.K., Cardinale B.J., Duffy J.E., Gamfeldt L. …, & Thompson P.L., A general biodiversity-function relationship is mediated by trophic level, Oikos, 2017, Vol. 126, pp. 18-31.

Onuchin A.A., Vlagooborot gornyh lesov Sibiri: Lokal’nye i regional’nye osobennosti, Dis. dokt. bil. nauk (Moisture rotation of mountain forests of Siberia: Local and regional features. Doctor’s thesis), Krasnoyarsk, 2003, 222 p.

Orlova M.A., Lukina N.V., Kamaev I.O., Smirnov V.E., Kravchenko T.V., Mozaichnost’ lesnyh biogeocenozov i produktivnost’ pochv (Forest ecosystem mosaics and soil fertility), Lesovedenie, 2011, No. 6, pp. 39-48.

Orlova M.A., Elementarnaya edinica lesnogo biogeocenoticheskogo pokrova dlya ocenki ekosistemnyh funkcij lesov (Elementary unit of the forest biogeocenotic cover for investigation of forest ecosystem functions), Trudy Karel’skogo nauchnogo centra. Seriya Ekologicheskie issledovaniya, 2013, No. 6, pp. 126-132.

Orlova M., Lukina N., Tutubalina O., Smirnov V., Isaeva G., Hofgaard F., Soil nutrient’s spatial variability in forest–tundra ecotones on the Kola Peninsula, Russia, Biogeochemistry, 2013, Vol. 113, pp. 283 -305.

Orlova M.A., Lukina N.V., Smirnov V.E., Artemkina N.A., Vliyanie eli na kislotnost’ i soderzhanie elementov pitaniya v pochvah severotaezhnyh el’nikov kustarnichkovo-zelenomoshnyh (The influence of spruce on acidity and nutrient content in soils of northern taiga dwarf shrub-green moss spruce forests), Pochvovedenie, 2016, No. 11, pp. 1355-1367.

Osipov A.F., Emissiya dioksida ugleroda s poverhnosti pochvy sosnyaka chernichno-sfagnovogo srednej tajgi (Carbon dioxide emission from the soil surface in a bilberry-sphagnum pine forest in the Middle Taiga), Pochvovedenie, 2013, No. 5, pp. 619-626.

Osipov A.F., Emissiya dioksida ugleroda s poverhnosti pochvy spelogo sosnyaka chernichnogo v srednej tajge Respubliki Komi (Carbon dioxide emission form the soil surface in mature bilberry pine forest in Middle Taiga of the Komi Republic), Lesovedenie, 2015, No. 5, pp. 356-366.

Osipov A.F., Bobkova K.S., Biologicheskaya produktivnost’ i fiksaciya ugleroda srednetaezhnymi sosnyakami pri perekhode iz srednevozrastnyh v spelye (Biological productivity and carbon sequestration of pine forests at transition from middle aged to mature in middle taiga), Lesovedenie, 2016, No. 5, pp. 346-354.

Osono T., Takeda H., Accumulation and release of nitrogen and phosphorus in relation to lignin decomposition in leaf litter of 14 tree species, Ecological Research, 2004, Vol. 19, No 6, pp. 593-602.

Pahomov A.E., Bulahov V.L., Bobylev Yu.P., Harakter, velichina i masshtaby royushchej deyatel’nosti krota v dolinnyh lesah stepnoj Ukrainy (The feature, magnitude and scale of burrowing activity of a mole in the valley forests of the steppe Ukraine), In: Ohrana i racional’noe ispol’zovanie zashchitnyh lesov stepnoj zony (Protection and rational use of protective forests of the steppe zone), Dnipropetrovsk, 1987, pp. 106-114.

Pahomov A.E., Formirovanie pochvennoj mezofauny pod vozdejstviem royushchih mlekopitayushchih v bajrachnyh dubravah Prisamar’ya (Soil Mesofauna Formation Effected by Mammalia Soil Burrowers in the Ravine Oak Forests of the Samara River Area), Vestnik zoologii, 2003, Vol. 37, No 1, pp. 41-48.

Paine R.T., The Pisaster‐Tegula interaction: Prey patches, predator food preference, and intertidal community structure, Ecology, 1969, Vol. 50, No 6, pp. 950-961.

Paustian K., Lehmann J., Ogle S., Reay D., Robertson G.P., Smith P., Climate-smart soils, Nature, 2016, Vol. 532, pp. 49-57.

Perel’ T.S., Rasprostranenie i zakonomernosti raspredeleniya dozhdevyh chervej fauny SSSR (Dispersal and patterns of spreading of earthworms of the fauna of the USSR), Moscow, Nauka, 1979, 272 p.

pleistocenepark.ru (December 12, 2020).

Poeydebat C., Jactel H., Moreira X., Koricheva J., Barsoum N., Bauhu, J., … & Gravel D., Climate affects neighbour-induced changes in leaf chemical defences and tree diversity-herbivory relationships, Functional Ecology, 2020, pp. 1-15.

Pogrebnyak P.S., Osnovy lesnoj tipologii (Fundamentals of forest typology), Kiev: AN USSR, 1955, 456 p.

Pollierer M.M., Scheu S., Tiunov A.V., Isotope analyses of amino acids in fungi and fungal feeding Diptera larvae allow differentiating ectomycorrhizal and saprotrophic fungi‐based food chains, Functional Ecology, 2020, Vol. 34, No 11, pp. 2375-2388.

Polyanskaya L.M., YUmakov D.D., Tyugaj Z.N., Stepanov A.L., Sootnoshenie gribov i bakterij v temnogumusovoj lesnoj pochve (Fungi and bacteria ratio in the dark humus forest soil), Pochvovedenie, 2020, No. 9, pp. 1094-1099.

Potapov A.M., Semenyuk I.I., Tiunov A.V., Seasonal and age-related changes in the stable isotope composition (15N/14N and 13C/12C) of millipedes and collembolans in a temperate forest soil, Pedobiologia, 2014, Vol. 57, No 4-6, pp. 215-222.

Potapov A.M., Tiunov A.V., Scheu S., Larsen T., Pollierer M.M., Combining bulk and amino acid stable isotope analyses to quantify trophic level and basal resources of detritivores: a case study on earthworms, Oecologia. 2019, Vol. 189, No 2, pp. 447-460.

Pretzsch H., Steckel M., Heym M., Biber P., Ammer C., Ehbrecht M., … & Vast F., Stand growth and structure of mixed-species and monospecific stands of Scots pine (Pinus sylvestris L.) and oak (Q. robur L., Quercus petraea (M att.) L iebl.) analysed along a productivity gradient through Europe, European Journal of Forest Research, 2020, Vol. 139, No 3, pp. 349-367.

Pugnaire F.I., Morillo J.A., Peñuelas J., Reich P.B., Bardgett R.D., Gaxiola A., … & Van Der Putten W.H., Climate change effects on plant-soil feedbacks and consequences for biodiversity and functioning of terrestrial ecosystems, Science advances, 2019, Vol. 5, No 11, p. eaaz1834.

Pukkala T., Instructions for optimal any-aged forestry, Forestry, An International Journal of Forest Research, 2018, Vol. 91, No 5, pp. 563-574.

Pulleman M.M., Six J., Uyl A., Marinissen J.C.Y., Jongmans A.G., Earthworms and management affect organic matterincorporation and microaggregate formation in agricultural soils, Appl. Soil Ecol., 2005, Vol. 29, pp. 1-15.

Pülzl H., Kleinschmit D., Arts B., Bioeconomy–an emerging meta-discourse affecting forest discourses? Scandinavian Journal of Forest Research, 2014, Vol. 29, No 4, pp. 386-393.

Punttila P., Kilpeläinen J., Distribution of mound-building ant species (Formica spp., Hymenoptera) in Finland: preliminary results of a national survey, Annales Zoologici Fennici, 2009, Vol. 46, No 1, pp. 1-15.

Rafes P.M., Rol’ i znachenie rastitel’noyadnyh nasekomyh v lesu (The role and significance of herbivorous insects in the forest), Moscow: Nauka, 1968, 233 p.

Ramenskij L.G., Vvedenie v kompleksnoe pochvenno-geobotanicheskoe issledovanie zemel’ (Introduction to integrated soil-geobotanical research of lands), Leningrad: Sel’hozgiz, 1938, 620 p.

Rämö J., Tahvonen O., Optimizing the harvest timing in continuous cover forestry, Environmental and Resource Economics, 2017, Vol. 67, pp. 853-868.

Rampino M.R., Shen S.Z., The end-Guadalupian (259.8 Ma) biodiversity crisis: the sixth major mass extinction? Historical Biology, 2019, pp. 1-7.

Rastitel’noyadnye zhivotnye v biogeocenozah sushi (Herbivorous animals in land biogeocenoses: Proc. All-Union Conference), Valdai, 3-6 June 1984, Moscow: Nauka, 1986, 189 p.

Reich P.B., Oleksyn J., Modrzynski J., Mrozinski P., Hobbie S.E., Eissenstat D.M., Tjoelker M.G., Linking litter calcium, earthworms and soil properties: a common garden test with 14 tree species, Ecology letters, 2005, Vol. 8, No 8, pp. 811-818.

Remezov N.P., Eshche o roli lesa v pochvoobrazovanii (More about the role of forests in pedogenesis), Pochvovedenie, 1956, No. 4, pp.70-79.

Remezov N.P., O roli lesa v pochvoobrazovanii (About the role of forests in pedogenesis), Pochvovedenie, 1953, No. 12, pp. 74-83.

Rooney T.P., Deer impacts on forest ecosystems: a North American perspective, Forestry: An International Journal of Forest Research, 2001, Vol. 74, No 3, pp. 201-208.

Roseberry J.L., Woolf A., Habitat-population density relationships for white-tailed deer in Illinois, Wildlife Society Bulletin, 1998, pp. 252-258.

Rosenfeld E.J., Assessing the ecological significance of linkage and connectivity for avian populations in urban areas, PhD thesis, University of Birmingham, 2012. 146 p.

Rozhkov A.C., Derevo i nasekomoe (Tree and insect), Novosibirsk: Nauka, 1981, 194 p.

Rubashko G.E., Hanina L.G., Smirnov V.E., Dinamika rastitel’nosti gruppirovok muravejnikov Formica rufa (Dynamics of plant communities related to the activity of Formica rufa ants), Zoologicheskij zhurnal, 2010, Vol. 89, No. 12, pp. 1448-1455.

Sablina T.B., Kopytnye Belovezhskoj Pushchi (Hoofed mammals of Belovezhskaya Pushcha), Moscow: Nauka, 1955, 192 p.

Saikkonen T., Vahtera V., Koponen S., Suominen O. Effects of reindeer grazing and recovery after cessation of grazing on the ground-dwelling spider assemblage in Finnish Lapland, PeerJ, 2019, Vol. 7, p. e7330.

Salemaa M., Derome J., Nojd P., Response of boreal forest vegetation to the fertility status of the organic layer along a climatic gradient, Boreal Environment Research, 2008, Vol. 13, pp. 48-66.

Sandor M., Schrader S., Earthworms affect mineralization of different organic amendments in a microcosm study, Bulletin of University of Agricultural Sciences and Veterinary Medicine Cluj-Napoca. Agriculture, 2007, Vol. 63, pp. 442-447.

Sauvadet M., Chauvat M., Brunet N., Bertrand I., Can changes in litter quality drive soil fauna structure and functions? Soil Biology and Biochemistry, 2017, Vol. 107, pp. 94-103.

Scharenbroch B.C., Bockheim J.G., Impacts of forest gaps on soil properties and processes in old growth northern hardwood-hemlock forests, Plant and soil, 2007, Vol. 294, No 1-2, pp. 219-233.

Scheller R.M., Mladenoff D.J., An ecological classification of forest landscape simulation models: tools and strategies for understanding broad-scale forested ecosystems, Landscape Ecology, 2007, Vol. 22, No 4, pp. 491-505.

Scherer-Lorenzen M., The functional role of biodiversity in the context of global change [in:] Forests and global change, D.A. Coomes, D.F.R.P. Burslem, W.D. Simonson (Eds.), Cambridge: Cambridge University Press, 2013, pp. 195-237.

Schlick-Steiner B.C., Steiner F.M., Moder K., Seifert B., Sanetra M., Dyreson E., … & Christian E.A., multidisciplinary approach reveals cryptic diversity in Western Palearctic Tetramorium ants (Hymenoptera: Formicidae), Molecular Phylogenetics and Evolution, 2006, Vol. 40, No 1, pp. 259-273.

Schliemann S.A., Bockheim J.G., Influence of gap size on carbon and nitrogen biogeochemical cycling in Northern hardwood forests of the Upper Peninsula, Michigan, Plant and soil, 2014, Vol. 377, No 1-2, pp. 323-335.

Schliemann S.A., Bockheim J.G., Methods for studying treefall gaps: a review, Forest ecology and management, 2011, Vol. 261, No 7, pp. 1143-1151.

Schmidt M.W., Torn M.S., Abiven S., Dittmar T., Guggenberger G., Janssens I. A., … & Nannipieri P., Persistence of soil organic matter as an ecosystem property, Nature, 2011, Vol. 478, No 7367, p. 49.

Schneider A.K., Hohenbrink T.L., Reck A., Zangerlé A., Schröder B., Zehe E., van Schaik L., Variability of earthworm-induced biopores and their hydrological effectiveness in space and time, Pedobiologia, 2018, Vol. 71, pp. 8-19.

Schuldt A., Assmann T., Brezzi M., Buscot F., Eichenberg D., Gutknecht J., … & Liu X., Biodiversity across trophic levels drives multifunctionality in highly diverse forests, Nature communications, 2018, Vol. 9, No 1, p. 2989.

Seidl R., Fernandes P.M., Fonseca T.F., Gillet F., Jönsson A.M., … & Mohren F., Modelling natural disturbances in forest ecosystems: a review, Ecological Modelling, 2011, Vol. 222, No 4, pp. 903-924.

Semenov M.V., Metabarkoding i metagenomika v pochvenno-ekologicheskih issledovaniyah: uspekhi, problemy i vozmozhnosti (Metabarcoding and metagenomics in soil ecology research: achievements, challenges and opportunities), ZHurnal obshchej biologii, 2019, Vol. 80, No. 6, pp. 403-417.

Shanin V.N., Komarov A.S., Mikhailov A.V., Bykhovets S.S., Modelling carbon and nitrogen dynamics in forest ecosystems of Central Russia under different climate change scenarios and forest management regimes, Ecological Modelling, 2011, Vol. 222, No 14, pp. 2262-2275.

Shanin V., Mäkipää R., Shashkov M., Ivanova N., Shestibratov K., Moskalenko S., … & Osipov A., New procedure for the simulation of belowground competition can improve the performance of forest simulation models, European Journal of Forest Research, 2015, Vol. 134, No 6, pp. 1055-1074.

Shanin V.N., Grabarnik P.YA., Byhovec S.S., Chertov O.G., Priputina I.V., Shashkov M.P., … & Ruchinskaya E.V., Parametrizaciya modeli produkcionnogo processa dlya dominiruyushchih vidov derev’ev Evropejskoj chasti RF v zadachah modelirovaniya dinamiki lesnyh ekosistem (Parametrization of the production process model for the dominant tree species in the European part of the Russian Federation in the problems of modeling the dynamics of forest ecosystems), Mathematical Biology and Bioinformatics, 2019, Vol. 14, No. 1, pp. 54-76.

Shanin V., Grabarnik P., Shashkov M., Ivanova N., Bykhovets S., FrolovP., Stamenov M., Crown asymmetry and niche segregation as an adaptation of trees to competition for light: conclusions from simulation experiments in mixed boreal stands, Mathematical and Computational Forestry and Natural-Resource Sciences, 2020, Vol. 12, No 1, pp. 26-49.

Shekhovtsov S.V., Rapoport I.B., Poluboyarova T.V., Geraskina A.P., Golovanova E.V., Peltek S.E., Morphotypes and genetic diversity of Dendrobaena schmidti (Lumbricidae, Annelida), Vavilov Journal of Genetics and Breeding, 2020, Vol. 24, No 1, pp. 48-54.

Shevchenko N.E., Rol’ Bison bonasus (Linnaeus, 1758) v formirovanii mozaiki prirodnogo lesnogo pokrova Vostochnoj Evropy. Soobshchenie pervoe. Dinamika areala i osobennosti troficheskoj i topicheskoj deyatel’nosti evropejskogo zubra v pozdnem golocene na territorii Vostochnoj Evropy (The role of Bison bonasus (Linnaeus, 1758) in the mosaic formation of natural forest cover in Eastern Europe. First article. The dynamics of the area, and features of the food and topical activity of the european bison in the Late Holocene in Eastern Europe), Russian Journal of Ecosystem Ecology, 2016, Vol. 1, No. 2, pp. 1-41.

Shevchenko N.E., Kuznecova A.I., Teben’kova D.N., Smirnov V.E., Geras’kina A.P., Gornov A.V., Tihonova E.V., Lukina N.V., Sukcessionnaya dinamika rastitel’nosti i zapasy pochvennogo ugleroda v hvojno-shirokolistvennyh lesah severo-zapadnogo Kavkaza (Succession dynamics of vegetation and storages of soil carbon in mixed forests of northwestern Caucasus), Lesovedenie, 2019, No. 3, pp. 163-176.

Siemann E., Carrillo J.A., Gabler C.A., Zipp R., Rogers W.E., Experimental test of the impacts of feral hogs on forest dynamics and processes in the southeastern US, Forest Ecology and Management, 2009, Vol. 248, pp. 546-533.

Simmons L.W., Ridsdill-Smith T.J. (eds.), Ecology and evolution of dung beetles, Oxford: Blackwell Publishing, 2011, pp. 1-20.

Six J., Bossuyt H., Degryze S., Denef K.A., History of research on the link between (micro) aggregates, soil biota, and soil organic matter dynamics, Soil and Tillage Research, 2004, Vol. 79, No 1, pp. 7-31.

Sklyarov G.A., K voprosu o deyatel’nosti krotov v pochvah dernovo-podzolistoj zony (To the question of the activity of moles in the soils of the sod-podzolic zone), Pochvovedenie, 1953, No. 8, pp. 51-57.

Skvorcova E.B., Ulanova N.G., Basevich V.F., Ekologicheskaya rol’ vetrovalov (The ecological role of windblows), Мoscow: Lesnaya promyshlennost’, 1983, 192 p.

Smirnov V.E., Funkcional’naya klassifikaciya rastenij metodami mnogomernoj statistiki (Functional classification of plants by multivariate analysis), Matematicheskaya biologiya i bioinformatika, 2007, Vol. 2, No. 1, pp. 1-17.

Smirnova O.V., Struktura travyanogo pokrova shirokolistvennyh lesov (Grass cover structure of broad-leaved forests), Moscow: Nauka, 1987, 207 p.

Smirnova O.V., Populyacionnaya organizaciya biocenoticheskogo pokrova lesnyh landshaftov (Population organization of biocenotic cover of forest landscapes), Uspekhi sovremennoj biologii, 1998, Vol. 118, No. 2, pp. 148-165.

Smirnova O.V., Hanina L.G., Smirnov V.E., Ekologo-cenoticheskie gruppy v rastitel’nom pokrove lesnogo poyasa Vostochnoj Evropy (Ecological-cenotic groups in the vegetation cover of forest belt of Eastern Europe), In: Vostochno-Evropejskie lesa: istoriya v golocene i sovremennost’ (Eastern European forests: history in Holocene and contemporaneity), Moscow: Nauka, 2004, Vol. 1, pp. 165-175.

Smirnova O.V., Prirodnaya organizaciya biogeocenoticheskogo pokrova lesnogo poyasa Vostochnoj Evropy. Teoreticheskie predstavleniya biogeocenologii i populyacionnoj biologii (The natural organization of biocenotic cover of forest belt of Eastern Europe. Theoretical concepts of biogeocenology and population biology), In: Vostochnoevropejskie lesa: istoriya v golocene i sovremennost’ (Eastern European forests: history in Holocene and contemporaneity), Moscow: Nauka, 2004, Vol. 1, pp. 14-25.

Smirnova O.V., Toropova N.A., Potential vegetation and potential ecosystem cover, Biology Bulletin Reviews, 2017, Vol. 7, No 2, pp. 139-149.

Smirnova O.V., Geraskina A.P., Korotkov V.N., Natural zonality of the forest belt of Northern Eurasia: myth or reality? Part 1 (literature review), Russian Journal of Ecosystem Ecology, 2020, Vol. 5, No 1, pp. 19-38.

Sobek S., Tscharntke T., Scherber C., Schiele S., Steffan-Dewenter I., Canopy vs. understory: Does tree diversity affect bee and wasp communities and their natural enemies across forest strata? Forest Ecology and Management, 2009, Vol. 258, No 5, pp. 609-615.

Söderbergh I., Ledermann T., Algorithms for simulating thinning and harvesting in five European individual-tree growth simulators: a review, Computers and Electronics in Agriculture, 2003, Vol. 39, No 2, pp. 115-140.

Solov’ev V.A., Biologiya i hozyajstvennoe znachenie barsukov Vyatsko-Kamskogo mezhdurech’ya, Dis. kand. biol. nauk (Biology and economic importance of badgers in the Vyatka-Kama interstream area, Candidate’s thesis,), Kirov, 2007, 162 p.

Spurgeon D.J. Keith A.M., Schmidt O., Lammertsma D.R., Faber J.H., Land-use and land-management change: relationships with earthworm and fungi communities and soil structural properties, BMC ecology, 2013, Vol. 13, No 1, p. 46.

Sredoobrazuyushchaya deyatel’nost’ zhivotnyh (Environment-forming activity of animals, Proc. Conf. Title), 17-18 December 1970, Moscow: Nauka, 1970, 101 p.

Steckel M., del Río M., Heym M., Aldea J., Bielak K., Brazaitis G., … & Jansons A., Species mixing reduces drought susceptibility of Scots pine (Pinus sylvestris L.) and oak (Quercus robur L., Quercus petraea (Matt.) Liebl.) – Site water supply and fertility modify the mixing effect, Forest Ecology and Management, 2020, Vol. 461, pp. 117908.

Storch D., Bohdalkovа E., Okie J., The more‐individuals hypothesis revisited: the role of community abundance in species richness regulation and the productivity–diversity relationship, Ecology Letters, 2018, Vol. 21, No 6, pp. 920-937.

Striganova B.R., Issledovanie roli mokric i dozhdevyh chervej v processah gumifikacii razlagayushchejsya drevesiny (Investigation of the role of woodlice and earthworms in the processes of humification of decaying wood), Pochvovedenie, 1968, No. 8, pp. 85-90.

Striganova B.R., Pitanie pochvennyh saprofagov (Nutrition of soil saprophages), Moscow: Nauka, 1980, 244 p.

Striganova B.R., Struktura i funkcii soobshchestv pochvoobitayushchih zhivotnyh (Structure and functions of communities of soil inhabiting animals), In: Strukturno-funkcional’naya rol’ pochv i pochvennoj bioty v biosphere (Structural and functional role of soils and soil biota in the biosphere), Moscow: Nauka, 2003, pp. 151-173.

Sukachev V.N., Rastitel’nye soobshchestva (Plant communities), Мoscow: Kniga, 1928, 232 p.

Sukachev V.N., Terminologiya osnovnyh ponyatij fitocenologii (Terminology of the basic concepts of phytocenology), Sovremennaya botanika, 1935, Vol. 5, pp. 11-21.

Sukachev V.N., Dinamika lesnyh biogeocenozov. Osnovy lesnoj biogeocenologii (Dynamics of forest biogeocenoses. Fundamentals of forest biogeocenology), Мoscow: Nauka, 1964, pp. 458-486.

Sukachev V.N., Osnovy lesnoj tipologii i biogeocenologii. Izbrannye trudy (Fundamentals of forest typology and biogeocenology. Selected Works), Leningrad: Nauka, 1972, 418 p.

Sundqvist M.K., Wardle D.A., Olofsson E., Giesler R., Gundale M.J., Chemical properties of plant litter in response to elevation: subarctic vegetation chalenges phenolic alocation theories, Functional Ecology, 2012, Vol. 26, No 3, pp. 1090-1099.

Suominen O., Olofsson J., Impacts of semi-domesticated reindeer on structure of tundra and forest communities in Fennoscandia: a review, Annales Zoologici Fennici, 2000, Vol. 37, No 4, pp. 233-249.

Swift M.J., Human impacts on biodiversity and ecosystem services: an overview [in:] The Fungal Community its Organization and Role in Ecosystems (Eds. J. Dighton, J.F. White, P. Oudemans), Boca Raton, FL: CRC Press, 2005, pp. 627-641.

Talashilkar S.C., Bhangarath P.P., Mehta V.B., Changes in chemical properties during composting of organic residues as influenced by earthworm activity, Journal of the Indian Society of Soil Science, 1999, Vol. 47, pp. 50-53.

The afterlife of a tree, Bobiec A. (Ed.), WWF Poland, 2005. 248 p.

The Economics of Ecosystems and Biodiversity: Mainstreaming the Economics of Nature. A synthesis of the approach, conclusions and recommendations of TEEB, Malta: Progress Press, 2010, 49 p.

Tihomirova L.G., O vliyanii royushchej deyatel’nosti krota na rastitel’nost’ lugov Moskovskoj oblasti (On the influence of the burrowing activity of a mole on the vegetation of meadows in the Moscow region), In: Struktura i funkcional’no-biogeocenoticheskaya rol’ zhivotnogo naseleniya sushi. Materialy soveshchaniya MOIP. Sekciya zoologii (The structure and functional-biogeocenotic role of the animal population of the land. Materials of the MOIP meeting. Zoology Section), Moscow: Nauka, 1967, pp. 97-99.

Tiunov A.V., Kuznecova N.A., Sredoobrazuyushchaya deyatel’nost’ nornyh dozhdevyh chervej (Lumbricus terrestris L.) i prostranstvennaya organizaciya pochvennoj bioty (Environmental activity of anecic earthworms (Lumbricus terrestris L.) and spatial organization of soil communities), Izvestiya Rossijskoj akademii nauk. Seriya biologicheskaya, 2000, No. 5, pp. 606-617.

Tiunov A.V., Metabioz v pochvennoj sisteme: vliyanie dozhdevyh chervej na strukturu i funkcionirovanie pochvennoj bioty, Avtoref. dis. dokt. biol. nauk (Metabiosis in soil system: impact of earthworms on the structure and functioning of soil biota, Extended abstract of Doctor’s thesis), Moscow: IPEE, 2007, 44 p.

Tobner C.M., Paquette A., Gravel D., Reich P.B., Williams L.J., Messier C., Functional identity is the main driver of diversity effects in young tree communities, Ecology letters, 2016, Vol. 19, No 6, pp. 638-647.

Tolkach V.N., Dvorak L.E., Izmenenie nadzemnoj fitomassy zhivogo napochvennogo pokrova pod vliyaniem dikih kopytnyh (Changes in the aboveground phytomass of the living ground cover under the influence of wild hoofed mammals), Belovezhskaya pushcha, 1980, Issue 4, pp. 29-38.

Toropova N.A., Rol’ geterotrofov v formirovanii mozaichno-yarusnoj struktury lesov (The role of heterotrophs in the formation of the mosaic-tiered structure of forests), In: Vostochnoevropejskie shirokolistvennye lesa (Eastern European broadleaf forests), Moscow: Nauka, 1994, pp. 228-241.

Tresch S., Frey D., Le Bayon R.C., Zanetta A., Rasche F., Fliessbach A., Moretti M., Litter decomposition driven by soil fauna, plant diversity and soil management in urban gardens, Science of the Total Environment, 2019, Vol. 658, pp. 1614-1629.

Treseder K.K., Lennon J.T., Fungal traits that drive ecosystem dynamics on land, Microbiology and Molecular Biology Reviews, 2015, Vol. 79, pp. 243-262.

Udra I.F., Rasselenie rastenij i voprosy paleo- i biogeografii (Plant dispersal and issues of paleo and biogeography), Kiev: AN USSR, 1988, 197 p.

Ulanova N.G., The effects of windthrow on forests at different spatial scales: a review, Forest ecology and management, 2000, Vol. 135, No 1-3. pp. 155-167.

Urban A.V., Prokushkin A.S., Korets M.A., Panov A.V., Gerbig C., Heimann M., Influence of the Underlying Surface on Greenhouse Gas Concentrations in the Atmosphere Over Central Siberia, Geography and Natural Resources, 2019, Vol. 40, No 3, pp. 221-229.

Van Breemen N., Finzi A.C., Plant-soil interactions: ecological aspects and evolutionary implications, Biogeochemistry, 1998, Vol. 42, pp. 1-19.

Van der Plas F., Biodiversity and ecosystem functioning in naturally assembled communities // Biological Reviews, 2019, Vol. 94, No 4, pp. 1220-1445.

Van der Plas F., Manning P., Allan E., Scherer-Lorenzen M., Verheyen K., Wirth C., … & Barbaro L. Jack-of-all-trades effects drive biodiversity–ecosystem multifunctionality relationships in European forests // Nature communications. 2016. Vol. 7. No. 1. P. 1-11.

Van der Plas F., Ratcliffe S., Ruiz‐Benito P., Scherer‐Lorenzen M., Verheyen K., Wirth C., … & Bastias C.C., Continental mapping of forest ecosystem functions reveals a high but unrealised potential for forest multifunctionality, Ecology letters, 2018, Vol. 21, No 1, pp. 31-42.

Van Groenigen J.W., Lubbers I.M., Vos H.M., Brown G.G., De Deyn G.B., Van Groenigen K.J., Earthworms increase plant production: a meta-analysis, Scientific report, 2014, Vol. 4, P. 63-65.

Van Groenigen J.W., Van Groenigen K.J., Koopmans G.F., Stokkermans L., Vos H.M., Lubbers I.M., How fertile are earthworm casts? A meta-analysis, Geoderma, 2019, Vol. 338, pp. 525-535.

Van Klink R., van Laar-Wiersma J., Vorst O., Smit C., Rewilding with large herbivores: Positive direct and delayed effects of carrion on plant and arthropod communities, PloS one, 2020, Vol. 15, No 1, p. e0226946.

Van Meerbeek K., Muys B., Schowanek S.D., Svenning J.C., Reconciling Conflicting Paradigms of Biodiversity Conservation: Human Intervention and Rewilding, BioScience, 2019, Vol. 69, No 12, pp. 997-1007.

Vasile M., The vulnerable bison: practices and meanings of rewilding in the Romanian Carpathians, Conservation and Society, 2018, Vol. 16, No 3, pp. 217-231.

Veen G.F.C., Olff H., Interactive effects of soil-dwelling ants, ant mounds and simulated grazing on local plant community composition, Basic and Applied Ecology, 2011, Vol. 12, No 8, pp. 703-712.

Vereshchagin N.K., Rusakov O.S., Kopytnye Severo-Zapada SSSR (istoriya, obraz zhizni i hozyajstvennoe ispol’zovanie (Hoofed mammals North-West of the USSR (history, way of life and practical use)), Leningrad: Nauka, 1979, 309 p.

Verheyen K., Vanhellemont M., Auge H., Baeten L., Baraloto C., Barsoum N., … & Haase J., Contributions of a global network of tree diversity experiments to sustainable forest plantations, Ambio, 2016, Vol. 45, No 1, pp. 29-41.

Vesterdal L., Clarke N., Sigurdsson B.D., Gundersen P., Do tree species influence soil carbon stocks in temperate and boreal forests? Forest Ecology and Management, 2013, Vol. 309, pp. 4-18.

Vicente-Silva J., Bergamin R. S., Zanini K.J.,   Pillar V.D.Mülle S.C., Assembly patterns and functional diversity of tree species in a successional gradient of Araucaria forest in Southern Brazil, Natureza & Conservação, 2016, Vol. 14, No 2, pp. 67-73.

Vostochnoevropejskie lesa: istoriya v golocene i sovremennost’ (Eastern European forests: history in the Holocene and contemporaneity), Moscow: Nauka, 2004, Vol. 1, 479 p.

Vostochnoevropejskie shirokolistvennye lesa (Eastern European broadleaf forests), Moscow: Nauka, 1994, 364 p.

Wagg C., Bender S.F., Widmer F., Van der Heijden M.G.A., Soil biodiversity and soil community composition determine ecosystem multifunctionality, Proceedings of the National Academy of Sciences, 2014, Vol. 11, No 14, pp. 5266-5270.

Wall D., Soil ecology and ecosystem services, Oxford, UK: Oxford University Press, 2012. 424 p.

Wardle D.A., Communities and Ecosystems: Linking Aboveground and Belowground Components, New Jerse: Princeton Univ. Press, Princeton, 2002. 391 p.

Whelan C.J., Şekercioğlu Ç.H., Wenny D.G., Why birds matter: from economic ornithology to ecosystem services, Journal of Ornithology, 2015, Vol. 156, No 1, pp. 227-238.

Wikström P., Edenius L., Elfving B., Eriksson L., Lämas T., Sonesson J., Öhman K., Wallerman J., Waller C., Klintebäck F., The Heureka Forestry Decision Support System: An Overview, Mathematical & Computational Forestry & Natural Resource Sciences, 2011, Vol. 3, No 2, pp. 87-95.

Wirthner S., The role of wild boar (Sus scrofa L.) rooting in forest ecosystems in Switzerland, A dissertation for the degree of doctor in science, Zurich, 2011, 103 p.

Wright J.P., Jones C.G., Flecker A.S., An ecosystem engineer, the beaver, increases species richness and the landscape scale, Oecologia, 2002, Vol. 132, pp. 96-101.

Wright J.P., Jones C.G. The concept of organisms as ecosystem engineers ten years on: progress, limitations, and challenges, BioScience, 2006, Vol. 56, No 3, pp. 203-209.

Yamamoto S.-I., Forest Gap Dynamics and Tree Regeneration, Journal of Forest Research, 2012, Vol. 5, No 4, pp. 223-229.

Yang X., Chen J., Plant litter quality influences the contribution of soil fauna to litter decomposition in humid tropical forests, southwestern China, Soil Biology and Biochemistry, 2009, Vol. 41, No 5, pp. 910-918.

Yatso K.N., Lilleskov E.A., Effects of tree leaf litter, deer fecal pellets, and soil properties on growth of an introduced earthworm (Lumbricus terrestris): implications for invasion dynamics, Soil Biology and Biochemistry, 2016, Vol. 94, pp. 181-190.

Yokoyama K., Kai H., Koga T., Kawaguchi S., Effect of dung beetle, Onthophagus lenzii H. on nitrogen transformation in cow dung and dung balls, Soil Science and Plant Nutrition, 1991, Vol. 37, No 2, pp. 341-345.

Zaharov A.A., Muravej, sem’ya, koloniya (Ant, family, colony), Moscow: Nauka, 1978, 144 p.

Zamolodchikov D.G., Korovin G.N., Utkin A.I., Chestnyh O.V., Songen B., Uglerod v lesnom fonde i sel’skohozyajstvennyh ugod’yah Rossii (Carbon in the forest fund and agricultural lands of Russia), Moscow: КМК, 2005, 212 p. 

Zaugol’nova L.B, Martynenko V.B., Opredelitel’ tipov lesa Evropejskoj Rossii (Guide on forest types in European Russia), 2012, Web-site, available at: http://www.cepl.rssi.ru/bio/forest/ (2020, 14 December).

Zaugol’nova L.B., Morozova O.V., Tipologija i klassifikacija lesov evropejskoj Rossii: metodicheskie podhody i vozmozhnosti ih realizacii (Typology and classification of European Russian forests: methodological approaches and potentialities of their realization), Lesovedenie, 2006, No. 1, pp. 34-48.

Zav’yalov N.A., Bobry (Castor fiber, C. canadensis) – sredoobrazovateli i fitofagi (Beavers (Castor fiber, C. canadensis) – founders of habitats and phytophages), Uspekhi sovremennoj biologii, 2013, Vol. 133, No. 5, pp. 502-528.

Zav’yalov N.A., Krylov A.V., Bobrov A.A., Ivanov V.K., Dgebuadze Yu.Yu., Vliyanie rechnogo bobra na ekosistemy malyh rek (Influence of river beaver on ecosystems of small rivers), Moscow: Nauka, 2005, 186 p.

Zav’yalova L.F., Biogeocenoticheckaya rol’ kabana v Darvinskom zapovednike i ego znachenie v sosednih sel’hozugod’yah (Biogeocenotic role of wild boar in the Darwin nature reserve and its importance in neighboring farmland), In: Nauchnye issledovaniya v zapovednikah i nacional’nyh parkah Rossii (federal’nyj otchet za 1992–1993 gody) (Scientific research in reserves and national parks of Russia (federal report 1992-1993)), Moscow, 1997, pp. 99-100.

Zenyakin S.A., Onipchenko V.G., Opyt ocenki masshtabov royushchej deyatel’nosti kavkazskogo krota (Talpa caucasica Satunin) na al’pijskom lugu Teberdinskogo zapovednika (Burrowing activity of the caucasian mole (Talpa caucasica Satunin) on an alpine meadow in the Teberda nature reserve), Byulleten’ MOIP. Otdel biologicheskij, 1997, Vol. 102, Issue 3, pp. 52-53.

Zhang D.Q., Hui D., Luo Y., Zhou G., Rates of litter decomposition in terrestrial ecosystems: global patterns and controlling factors, Journal of Plant Ecology, 2008, Vol. 1, No 2, pp. 85-93.

Zhang C., Mora P., Dai J., Chen X., Giusti-Miller S., Ruiz-Camach N., … & Lavelle P., Earthworm and organic amendment effects on microbial activities and metal availability in a contaminated soil from China, Applied Soil Ecology, 2016, Vol. 104, pp. 54-66.

Zhang B., Lu X., Jiang J., DeAngelis D.L., Fu Z., Zhang J., Similarity of plant functional traits and aggregation pattern in a subtropical forest, Ecology and Evolution, 2017, Vol. 7, No 12, pp. 4086-4098.

Zhu J., Lu D., Zhang W., Effects of gaps on regeneration of woody plants: a meta-analysis, Journal of Forestry Research, 2014, Vol. 25, No 3, pp. 501-510.

Zimov S.A., Pleistocene park: return of the mammoths ecosystem, Science, 2005, Vol. 308, No 5723, pp. 796-798.

Zimov S.A., Zimov N.S., Tikhonov A.N., Chapin III F.S., Mammoth steppe: a high-productivity phenomenon, Quaternary Science Reviews, 2012, Vol. 57, pp. 26-45.

Zlotin R.I. Hodasheva K.I., Rol’ zhivotnyh v biologicheskom krugovorote lesostepnyh ekosistem (The role of animals in the biological cycle of forest-steppe ecosystems), Moscow: Nauka, 1974, 217 p.

Zryanin V.A., Vliyanie murav’ev roda Lasius na pochvy lugovyh biogeocenozov (Effects of ants of the genus Lasius on soils of meadow biogeocenoses), Uspekhi sovremennoj biologii, 2003, Vol. 123, No. 3, pp. 278-287.