• DOI: 10.31509/2658-607x-202583-173
  • УДК 57. 574.4

SOIL ORGANOPEDOGENESIS AS AN UNHEEDED BASIC SOIL FORMING PROCESS

O. G. Chertov*, N. V. Lukina

Isaev Centre for Forest Ecology and Productivity of the RAS

Profsoyuznaya st. 84/32 bldg. 14, Moscow, 117997, Russia

*E-mail: ochertov@rambler.ru

Received: 21.05.2025

Revised: 01.07.2025

Accepted: 15.07.2025

A concept of soil organopedogenesis has been proposed as an unheeded active process of general pedogenesis reflecting a predominant significance of biological factor in soil formation according to the Vernadsky’s theory on organism’s leading role in all natural processes of continental part of the biosphere. A methodology for the including organopedogenesis in the modern soil classifications has been proposed as well. It is the integration into existing Russian soil taxonomy system the types of soil organic matter accumulation and transformation (humus forms) at the level with the types of organoprofiles. The inclusion of these components will make the classification dynamic and increase its practical importance that is especially significant for decision making at the time being with the fast negative global environmental changes.

 Keywords: biological factor of soil formation, organoprofile, SOM accumulation and transformation, humus forms

 REFERENCES

Abakumov E. V., Hronologiya ontogeneza pervichnyh pochv: obzor problemy (Chronology of ontogenesis of primary soils: a review of the problem), Vestnik SPbGU, Ser. 3, 2011, Vyp. 3, pp. 114–120.

Abakumov E. V., Soils of mining areas – natural model of soil formation, Proceed. 1st Intern. Conf. on Soils and Archaeology, Szazhalombatta, Hungary, 2001, pp. 163−165.

Alekseev V. A., Lesnye ekosistemy i atmosfernoe zagryaznenie (Forest ecosystems and atmospheric pollution), Leningrad: Nauka 1990, 200 p.

Androhanov V. A., Ovsyannikova S. V., Kurachev V. M., Technozems: properties, regimes, functioning (Technosols: properties, modes, functioning), Novosibirsk: Nauka, 2000, 200 p.

Bahmet O. N., Strukturno-funkcional’naya organizaciya organoprofilej pochv lesnyh ekosistem Severo-Zapada Rossii (Structural and functional organization of soil organoprofiles of forest ecosystems in Northwest Russia), Avtoref. diss. dokt. biol. nauk (Abst. diss. doct. biol. sci.), Petrozavodsk, 2015, 49 p.

Bioraznoobrazie i funkcionirovanie lesnyh ekosistem (Biodiversity and functioning of forest ecosystems), N. V. Lukina (ed.), Moscow: KMK, 2021, 327 p.

Blagovidov N. L., Burkov G. L., Metodicheskie ukazaniya k proizvodstvu pochvennyh issledovanij i harakteristike uslovij mestoobitaniya lesa (Methodical instructions for the production of soil studies and characterization of forest habitat conditions), Leningrad: LTA, 1959. 30 p.

Bobrovskij M. V., Lesnye pochvy evropejskoj Rossii. Bioticheskie i antropogennye faktory formirovaniya (Forest soils of European Russia. Biotic and anthropogenic factors of formation), Moscow: KMK, 2010, 392 p.

Bogatyryov L. G., Demin V. V., Matyshak G. V., Sapozhnikova V. A., O nekotoryh teoreticheskih aspektah issledovaniya lesnyh podstilok (On some theoretical aspects of forest litter research), Lesovedenie, 2004, No 4, pp. 17–29.

Bogatyryov L. G., Fomina T. V., Harakteristika podstilok sosnyakov Prisurskogo lesnogo massiva (Characteristics of pine forest’s litters of the Prisursky forest massif), Vestnik Moskovskogo universiteta, Seriya 17, Pochvovedenie, 91, No 3, pp. 28–39.

Broll G., Brauckmann H.-J., Overesch M., Junge B., Erber C., Milbert, G., Baize D., Nachtergaele F., Topsoil characterization – Recommendations for revision and expansion of the FAO-Draft (1998) with emphasis on humus forms and biological features, Journal of Plant Nutrition and Soil Science. 2006, Vol. 169, No 3, pp. 453–461.

Budyko M. I., Evolyuciya biosfery (Evolution of the biosphere), Leningrad: Gidrometeoizdat, 1984, 488 p.

Chertov O. G. O smyslovoj strukture naimenovanij taksonov pochvennyh klassifikacij (On the semantic structure soil classification taxa names), Voprosy lesnoj nauki, 2025, Vol. 8, No 1, Article 163, DOI: 10.31509/2658-607x-202581-163

Chertov O. G., Dyrenkov S. A., O klassifikacii lesnyh biogeocenozov Karel’skogo pereshejka (On the classification of forest biogeocenoses of the Karelian Isthmus), Trudy Petrozavodskoj lesnoj opytnoj stacii, 1973, Vol. 2, pp. 74–80.

Chertov O. G., Ekologiya lesnyh zemel’. Pochvenno-ekologicheskoe issledovanie lesnyh mestoobitanij (Ecology of forest lands. Soil-ecological study of forest sites), Leningrad: Nauka, 1981, 192 p.

Chertov O. G., Ekotopy tropicheskogo dozhdevogo lesa (Ecotopes of tropical rain forest), Leningrad: Nauka, 1985, 47 p.

Chertov O. G., Filippov G. V., Mel’nickaya G. B., Krupnomasshtabnoe kartirovanie pochv pri lesoustrojstve, Metodicheskie ukazaniya (Large-scale mapping of soils in forest management. Methodological Instructions), Leningrad: Len. NII lesn. hoz-va, 1978, 52 p.

Chertov O. G., Mel’nickaya G. B., Lamov A. K., Berg I. E., Opyt rabot po lesnoj tipologii na osnove kartirovaniya pochv i mestoobitanij (Experience of work on forest typology based on mapping of soils and sites), Sbornik statej po lesovodstvu, Leningrad: Len. NII lesn. hoz-va, 1974, No 17, pp. 24–47.

 Chertov O. G., Nadporozhskaya M. A., Development and application of humus form concept for soil classification, mapping and dynamic modelling in Russia, Applied Soil Ecology, 2018, Vol. 122, pp. 420–423, DOI: 10.1016/j.apsoil.2017.04.006

Chertov O. G., Nadporozhskaya M. A., Formy gumusa lesnyh pochv: koncepcii i klassifikacii (Humus forms of forest soils: concepts and classifications), Pochvovedenie, 2018, No 10, pp. 1202–1214.

Chertov O. G., Nadporozhskaya M. A., Palyonova M. M., Priputina I. V., Edafologiya v strukture pochvovedeniya i ekosistemnoj ekologii (Edaphology in the structure of soil science and ecosystem ecology), Russian Journal of Ecosystem Ecology, 2018, Vol. 3, No 3, DOI: 10.21685/2500-0578-2018-3-2

Chertov O. G., O ekologicheskih funkciyah i evolyucii pochv (On ecological functions and evolution of soils), Vestnik LGU, Ser. 3, 1990, No 2(10), pp. 75–81.

Chertov O. G., Opredelenie tipov gumusa lesnyh pochv (Definition of humus types in forest soils), Metodicheskie ukazaniya, Leningrad: Len. NII lesn. hoz-va, 1974, 16 p.

Chertov O. G., Razumovskij S. M., Ob ekologicheskoj napravlennosti processov razvitiya pochv (On the ecological orientation of soil development processes), Zhurnal obshchej biologii, 1980, Vol. 62, No 3, pp. 386–396.

Chertov O., Gorbushina A., Deventer B., A model for microcolonial fungi growth on rock surfaces, Ecological Modelling, 2004, Vol. 177, No 3-4, pp. 415–426, DOI: 10.1016/j.ecolmodel.2004.02.011

Chukov S. N., Primenenie metoda yadernogo magnitnogo rezonansa k izucheniyu organicheskogo veshchestva pochv (Application of the nuclear magnetic resonance method to the study of soil organic matter), Vestnik Sankt-Peterburgskogo universiteta. Seriya biologicheskaya, 1997, No 4, pp. 50–59.

Danilov D. A., Yakovlev A. A., Zajcev D. A., Ivanov A. A., Izmeneniya v postagrogennyh pochvah v hode vosstanovleniya drevesnoj rastitel’nosti v usloviyah yugo-zapada Leningradskoj oblasti (Changes in postagrogenic soils during the restoration of woody vegetation in the southwest of the Leningrad region), 2024, Izvestiya SPbLTA, No 251, DOI: 10.21266/2079-4304.2024.251.97-122

Dobrovol’skij G. V., Karpachevskij L. O., Kriksunov E. A., Geosfery i pedosfera (Geospheres and pedosphere), Moscow, GEOS, 2010, 190 p.

Dobrovol’skij G. V., Pedosfera – obolochka zhizni planety Zemlya (The pedosphere is the shell of life on planet Earth), Biosfera, 2009, Vol. 1, No 1, pp. 6–14.

Dokuchaev V. V., Russkij chernozyom: Otchyot Vol’nomu ekonomicheskomu obshchestvu (Russian chernozem: Report to the Free Economic Society), St. Petersburg: Tip. Deklerona i Evdokimova, 1883, 376 p.

 Duchaufour Ph., Precis de Pedologie, 2nd edition, Paris: Masson, 1965, 482 p.

Dyrenkov S. A., Struktura i dinamika taezhnyh el’nikov (Structure and dynamics of taiga spruce forests), Leningrad: Nauka, 1984, 176 p.

Dyushofur F., Osnovy pochvovedeniya. Evolyuciya pochv (Fundamentals of Soil Science. Soil Evolution). Moscow: Progress, 1979, 592 p.

 Emmer I. M., Humus form and soil development during a primary succession of monoculture Pinus sylvestris forests on poor sandy substrates, Amsterdam, 1995, 135 p.

Evolyuciya pochv i pochvennogo pokrova. Teoriya, raznoobrazie prirodnoj evolyucii i antropogennyh transformacij pochv (Evolution of soils and soil cover. Theory, diversity of natural evolution and anthropogenic transformations of soils), V. N. Kudeyarov, I. V. Ivanov (eds.), Moscow: GEOS, 2015, 928 p.

Evolyuciya, funkcionirovanie i ekologicheskaya rol’ pochv kak komponentov biosfery (Evolution, functioning and ecological role of soils as components of the biosphere), A. O. Alekseev (ed.), Moscow: KMK, 2020, 290 p.

Fedorchuk V. N., Neshataev V. Yu., Kuznecova M. L., Lesnye ekosistemy severo-zapadnyh rajonov Rossii. Tipologiya, dinamika, hozyajstvennye osobennosti (Forest ecosystems of northwestern Russia. Typology, dynamics, and economic characteristics), O. G. Chertov (ed.), St. Petersburg: St. Petersburg Research Institute of Forestry, 2005, 381 p.

Fokin A. D., Dinamicheskaya harakteristika gumusovogo profilya podzolistoj pochvy (Dynamic characteristics of the humus profile of podzolic soil), Izvestiya Timiryazevskoj sel’skohozyajstvennoj akademii, 1975, No 4, pp. 80–88.

Fridland V.  M., Pochvy i kory vyvetrivaniya vlazhnyh tropikov (Soils and weathering crusts of the humid tropics), AN SSSR, Pochvennyj in-t im. V. V. Dokuchaeva, Moscow: Nauka, 1964, 312 p.

Gerasimova M. I., Nuzhen li uroven’ roda v klassifikacii pochv Rossii? (Is level ROD (genus) nessesary in Russian soil classification?) Byulleten’ Pochvennogo instituta im. V. V. Dokuchaeva, 2018, Vyp. 95, pp. 90–98, DOI: 10.19047/0136-1694-2018-95-90-98

Glazovskaya M. A., Geohimiya prirodnyh i tekhnogennyh landshaftov SSSR (Geochemistry of natural and technogenic landscapes of the USSR), Moscow: Vysshaya shkola, 1988. 327 p.

Gorshkov V. G., Fizicheskie i biologicheskie osnovy ustojchivosti zhizni (Physical and biological foundations of life sustainability), Moscow, 1995, 470 p.

Green R. N., Trowbridge R. L., Klinka K., Towards a taxonomic classification of humus forms, Forest Science, 1993, Vol. 39, No 1, pp. 1–56.

Grigor’ev G. I., Diagnosticheskie pokazateli stepeni okul’turennosti dernovo-podzolistyh, svetlo-seryh i seryh lesnyh suglinistyh pochv (Diagnostic indicators of the degree of agronomic improvement of sod-podzolic, light gray and gray forest loamy soils), Teoreticheskie osnovy i metody opredeleniya optimal’nyh parametrovsvojstv pochv, Moscow, 1980, pp. 21–29.

Grishina L. A., Gumusoobrazovanie i gumusnoe sostoyanie pochv (Humus formation and humus state of soils), Moscow: MGU, 1986, 244 p.

Grishina L. A., Orlov D. S., Sistema pokazatelej gumusnogo sostoyaniya pochv (System of indicators of humus status of soils), Problemy pochvovedeniya (Sovetskie pochvovedy k II Mezhdunarodnomu kongressu pochvovedov v Kanade, 1978), Moscow: 1978, pp. 42–47.

Hitrov N. B., Gerasimova M. I., Predlagaemye izmeneniya v klassifikacii pochv Rossii: diagnosticheskie priznaki i pochvooobrazuyushchie porody (Proposed changes in the classification of soils of Russia: diagnostic features and parent rocks), Pochvovedenie, 2022, No 1, pp. 3–14, DOI: 10.31857/S0032180X22010087

 Humusica – Terrestrial Natural Humipedons, A. Zanella, J. Ascher-Jenull (eds.), Applied Soil Ecology, 2018, Vol. 122, Part I, II, III.

IUSS Working Group WRB. World Reference Base for Soil Resources. International soil classification system for naming soils and creating legends for soil maps, 4th edition, International Union of Soil Sciences (IUSS), Vienna, Austria, 2022, DOI: 10.29003/m4174.978-5-317-07235-3.

Kalinina O., Chertov O., Nadporozhskaya M., Giani L., Properties of soil organic matter of Plaggic Anthrosols from Northwest Germany, Northwest and North Russia, Archives of Agronomy and Soil Science, 2009, Vol. 55, No 5, pp. 477–492, DOI: 10.1080/03650340802637733

Karpachevskij L. O., Les i lesnye pochvy (Forest and forest soils), Moscow: Lesnaya promyshlennost’, 1981, 264 p.

Karpachevskij L. O., Stroganova M. P., Targul’yan V. O., Vasenev I. I., Goncharuk N. Yu., Pochvennye mikrosukcessii v klimaksnyh el’nikah yuzhnoj tajgi (Soil microsuccessions in climax spruce forests of the southern taiga, Istoriya razvitiya pochv SSSR v golocene (History of soil development in the USSR in the Holocene), Pushchino: AN SSSR, 1984, pp. 68–69.

Klassifikaciya i diagnostika pochv Rossii (Classification and diagnostics of soils in Russia), L. L. Shishov, V. D. Tonkonogov, I. I. Lebedeva, M. I. Gerasimova (compl.), Smolensk: Ojkumena, 2004, 342 p.

Klinka K., Krestov P., Fons J., Chourmouzis C., Towards a taxonomic classification of humus forms: Third approximation, Scientia Silvia, 1997, No 9, pp. 1–5.

Kõlli R., Köster T., Interrelationships of humus cover (pro humus form) with soil cover and plant cover: humus form as transitional space between soil and plant, Applied Soil Ecology, 2018, Vol. 123, pp. 451–454.

Komarov A., Chertov O., Bykhovets S., Shaw C., Nadporozhskaya M., Frolov P., Shashkov M., Shanin V., Grabarnik P., Priputina I., Zubkova E., Romul_Hum model of soil organic matter formation coupled with soil biota activity. I. Problem formulation, model description, and testing, Ecological Modelling, 2017, Vol. 345, pp. 113–124, DOI: 10.1016/j.ecolmodel.2016.08.007

Kostychev P.  A., Pochvy chernozemnoj oblasti Rossii, ih proiskhozhdenie, sostav i svojstva (Soils of the chernozem region of Russia, their origin, composition and properties), St. Petersburg: Izd. Devriena, 1886, 231 p.

Kovda V.  A., Biogeohimiya pochvennogo pokrova (Biogeochemistry of soil cover), Moscow, 1985, 493 p.

Kruedener A. A., Osnovy klassifikacii tipov nasazhdenij. Materialy po izucheniyu russkogo lesa (Basics of classification of stand types. Materials on the study of the Russian forest), Petrograd, 1916, 190 p.

Kubiena W. L., The soils of Europe. London: Thomas Murby Co., 1953, 318 p.

Lebedeva H. H., Tonkonogov V. D., Gerasimova M. I., Opyt razrabotki faktornoj klassifikacii pochv (Experience in developing a factor classification of soils), Pochvovedenie, 2000, No 2, pp. 148–157.

Lehmann J., Kleber M., The Contentious Nature of Soil Organic Matter, Nature, 2015, Vol. 528, pp. 60–68, DOI: 10.1038/nature16069

Lessovaia S., Chertov O., Goryachkin S., Specificity of pedogenesis in shallow soils on massive rocks of East Fennoscandia, Agricultural Sciences/Zemès ukio Mokslai, 2008, Vol. 12, No 3, pp. 80–86.

Lukina N. V., Bartalev S. A., Geras’kina A. P., Plotnikova A. S., Gornov A. V., Ershov D. V., … & Ruchinskaya E. V., Rol’ starovozrastnyh lesov v akkumulyacii i hranenii ugleroda (The role of old-growth forests in carbon accumulation and storage), Izvestiya Rossijskoj akademii nauk. Seriya geograficheskaya, 2023, Vol. 87, No 4, рр. 536–557.

Lukina N. V., Geraskina A. P., Kuznecova A. I., Smirnov V. E., Gornov A. V., Shevchenko N. E., Tihonova E. V., Funkcional’naya klassifikaciya lesov: aktual’nost’ i podhody k razrabotke (Functional classification of forests: relevance and approaches to development), Lesovedenie, 2021, No 6, pp. 566–580.

Lukina N. V., Kuznecova A. I., Geraskina A. P., Smirnov V. E., Ivanova V. N., Teben’kova D. N., Gornov A. V., Shevchenko N. E., Tihonova E. V., Neuchtennye faktory, opredelyayushchie zapasy ugleroda v lesnyh pochvah, Meteorologiya i gidrologiya (Unaccounted factors determining carbon reserves in forest soils), 2022, No 10, pp. 92–110.

Lukina N. V., Nikonov V. V., Biogeohimicheskie cikly v lesah Severa v usloviyah aerotekhnogennogo zagryazneniya (Biogeochemical cycles in northern forests under conditions of aerotechnogenic pollution), Apatity: Kol’skij nauchnyj centr RAN, 1996, 192 р.

Lukina N. V., Nikonov V. V., Pitatel’nyj rezhim lesov severnoj tajgi: prirodnye i tekhnogennye aspekty (Nutrient regime of northern taiga forests: natural and technogenic aspects), Apatity: Kol’skij nauchnyj centr RAN, 1998, 315 р.

Lukina N. V., Orlova M. A., Bahmet O. N., Tihonova E. V., Teben’kova D. N., Kazakova A. I., Kryshen’ A. M., Gornov A. V., Smirnov V. E., Shashkov M. P., Ershov V. V., Knyazeva S. V., Vliyanie rastitel’nosti na harakteristiki lesnyh pochv Respubliki Kareliya (Influence of vegetation on the characteristics of forest soils of the Republic of Karelia), Pochvovedenie, 2019, No 7, pp. 827–842.

Lukina N. V., Orlova M. A., Isaeva L. G., Plodorodie lesnyh pochv kak osnova vzaimosvyazi pochva-rastitel’nost’ (Fertility of forest soils as a basis for soil-vegetation relationships), Lesovedenie, 2010, No 5, pp. 45–56.

Lukina N. V., Polyanskaya L. M., Orlova M. A., Pitatel’nyj rezhim pochv severotaezhnyh lesov (Nutrient regime of soils of northern taiga forests), Moscow: Nauka, 2008, 341 р.

Lukina N. V., Tikhonova E. V., Danilova M. A., Bakhmet O. N., Kryshen A. M., …, & Zukert N. V., Associations between forest vegetation and the fertility of soil organic horizons in northwestern Russia, Forest Ecosystems, 2019, pp. 1–19, doi: 10.1186/s40663-019-0190-2

Lukina N. V., Vliyanie sovremennyh izmenenij klimata na lesnye pochvy (The impact of modern climate change on forest soils), Seminar Instituta global’nogo klimata i ekologii imeni akademika Yu. A. Izraelya, IGKE, 2025, URL: clck.ru/3PJV2o (2025, 01 July).

Mahonina G. I., Nachal’nye processy pochvoobrazovaniya v tekhnogennyh ekosistemah Urala (Initial processes of soil formation in technogenic ecosystems of the Urals), Avtoref. diss. dokt. biol. nauk (Abst. diss. doct. biol. sci.), Tomsk, 2004, 38 p.

Malysheva N. V., Filipchuk A. N., Zolina T. A., Kinigopulo P. S., SHalimova E. M., Popik S. A., Sil’nyagina G. V., Analiz zarubezhnogo opyta nacional’nyh inventarizacij lesov: metody, vyborka, rezul’taty i mezhdunarodnaya statistika (Analysis of foreign experience of national forest inventories: methods, sampling, results and international statistics), Lesohozyajstvennaya informaciya, 2022, No 2, pp. 90–132, DOI: 10.24419/LHI.2304-3083.2022.2.08

Min’kovskij G. M., Shoba S. A., Morfologiya i klassifikaciya organoprofilej pochv yuzhnoj tajgi (Morphology and classification of soil organoprofiles in the southern taiga), Pochvovedenie, 1994, No 9, pp. 90–101.

Min’kovskij G. M., Shoba S. A., Morfostrukturnye podhody k tipizacii organoprofilej pochv (Morphostructural approaches to the typology of soil organoprofiles), Pochvovedenie, 1995, No 10, pp. 1271–1283.

Mirovaya referativnaya baza pochvennyh resursov. Mezhdunarodnaya sistema pochvennoj klassifikacii dlya diagnostiki pochv i sostavleniya legend pochvennyh kart (World database of soil resources. International system of soil classification for soil diagnostics and compilation of soil map legends: translate from English), M. I. Gerasimova, P. V. Krasil’nikov (eds.), 4th edition, Mezhdunarodnyj soyuz nauk o pochve, Moscow: MAKS Press, 2024, 248 p.

Müller P. E., Studien über die natürlichen Humusformen und deren Entwicklung auf Vegetation und Boden, Berlin, 1887, 324 p.

Nadporozhskaya M. A., Mohren G. M. J., Chertov O. G., Komarov A. S., Mikhailov A. V., Dynamics of soil organic matter in primary and secondary forest succession on sandy soils in The Netherlands: An application of the ROMUL model, Ecological Modelling, 2006, Vol. 190, pp. 399–418, DOI: 10.1016/j.ecolmodel.2005.03.025

Odum Yu., Ekologiya (Ecology), Vol. 2, Moscow, Mir, 1986, 376 p.

Orlov D. S., Biryukova O. N., Suhanova N. I., Organicheskoe veshchestvo pochv Rossijskoj Federacii (Organic matter of soils of the Russian Federation), Moscow: Nauka, 1996, 256 p.

Perel’man A. I., Geohimiya landshafta (Geochemistry of the landscape), Moscow: Vysshaya shkola, 1975, 341 p.

Piccolo A., Spaccini R., Savy D., Drosos M., Cozzolino V., The Soil Humeome: Chemical structure, functions and technological perspectives, In: Sustainable Agrochemistry / Vaz Jr. Silvio (ed.), Springer, Cham, 2019, DOI: 10.1007/978-3-030-17891-8_7

Polevoj opredelitel’ pochv (Field guide to soils), Moscow: Pochvennyj in-t im. V. V. Dokuchaeva, 2008, 182 p.

Ponge J.-F., Humus forms in terrestrial ecosystems: a framework to biodiversity, Soil Biology and Biochemistry, 2003, Vol. 35, No 7, pp. 935–945, DOI: 10.1016/s0038-0717(03)00149-4.hal-00498465

Ponomareva V. V., Teoriya podzoloobrazovatel’nogo processa (Theory of the podzol-forming process), Moscow-Leningrad: Nauka, 1964, 379 р.

Prokashev A. M., Pochvy so slozhnym organoprofilem yuga Kirovskoj oblasti (Soils with a complex organoprofile in the south of the Kirov region), Kirov, 1999, 176 p.

Razumovskij S. M., Izbrannye Trudy (Selected works), Moscow: KMK, 1999, 560 р.

Reintam L., Elmar K., Rooma I., Development of soil organic matter under pine on quarry detritus of open-cast oil-shale mining, Forest Ecology and Management, 2002, Vol. 171, pp. 191−198.

Rode A. A., Genezis pochv i sovremennye faktory pochvoobrazovaniya (Soil genesis and modern factors of soil formation), Moscow: Nauka, 1984, 256 p.

Rodin L. E., Bazilevich N. I., Dinamika organicheskogo veshchestva i biologicheskij krugovorot zol’nyh elementov i azota v osnovnyh tipah rastitel’nosti zemnogo shara (Dynamics of organic matter and biological cycle of ash elements and nitrogen in the main vegetation types of the globe), Akad. nauk SSSR. Botan. in-t im. V. L. Komarova, Moscow-Leningrad: Nauka, 1965, 253 p.

Semenov V. M, Kogut B. M., Pochvennoe organicheskoe veshchestvo (Soil Organic Matter), Moscow: GEOS, 2015, 233 p.

Semenov V. M., Lebedeva T. N., Sokolov D. A., Zinyakova N. B., Lopes de Gerenyu V. O., Semenov M. V., Izmerenie pochvennyh pulov organicheskogo ugleroda, vydelennyh bio-fiziko-himicheskimi sposobami frakcionirovaniya (Measurement of the soil organic carbon pools isolated using bio-physical-chemical fractionation methods), Pochvovedenie, 2023, No 9, pp. 1155–1172, DOI: 10.31857/S0032180X23600427

Shanin V. N., Frolov P. V., Priputina I. V., Chertov O. G., Byhovec S. S., Zubkova E. V., Portnov A. M., Frolova G. G., Stamenov M. N., Grabarnik P. Ya., Modelirovanie dinamiki lesnyh ekosistem s uchyotom ih strukturnoj neodnorodnosti na raznyh funkcional’nyh i prostranstvennyh urovnyah (Modeling the dynamics of forest ecosystems taking into account their structural heterogeneity at different functional and spatial levels), Voprosy lesnoj nauki, 2022, Vol. 6, No 3, DOI: 10.31509/2658-607x-202252-112

Shoba S. A., Morfogenez pochv lesnoj zony (Morphogenesis of soils in the forest zone), Avtoref. diss. dokt. biol. nauk (Abst. diss. doct. biol. sci.), Moscow, 1988, 48 p.

Shumakov B. C., Principy klassifikacii, nomenklatury i kartirovaniya lesnyh podstilok (Principles of classification, nomenclature and mapping of forest litter), Sbornik rabot po lesnomu hozyajstvu, Vyp. 35, Moscow–Leningrad: Goslesbumizdat, 1958, рр. 199–216.

Sibircev N. M., Pochvovedenie (Soil Science), St. Petersburg: Izd. Skvorcova, 1899, 360 p.

Smirnova O. V., Populyacionnaya organizaciya biocenoticheskogo pokrova lesnyh landshaftov (Population organization of the biocenotic cover of forest landscapes), Uspekhi sovremennoj biologii, 1998, Vol. 118, No 2, pp. 148–165.

Smirnova O. V., Toropova N. A., Potencial’naya rastitel’nost’ i potencial’nyj ekosistemnyj pokrov (Potential vegetation and potential ecosystem cover), Uspekhi sovremennoj biologii, 2016, Vol. 136, No 2, pp. 199–211.

Smirnova O. V., Toropova N. A., Sukcessiya i klimaks kak ekosistemnyj process (Succession and climax as an ecosystem process), Uspekhi sovremennoj biologii, 2008, Vol. 128, No 2, pp. 129–144.

Sokolov I. A., Pochvoobrazovanie i ekzogenez (Soil formation and exogenesis), Moscow, 1997, 244 p.

Solodovnikov A. N., Modeli nachal’nogo razlozheniya hvoi sosny obyknovennoj v srednetaezhnyh usloviyah Vostochnoj Fennoskandii (Models of initial decomposition of Scots pine needles in the middle taiga conditions of Eastern Fennoscandia), Byulleten’ Pochvennogo instituta im. V. V. Dokuchaeva, 2025, Vyp, 122, pp. 41–61, DOI: 10.19047/0136-1694-2025-122-41-61

Sukachev V. N., Dylis N. V., Osnovy lesnoj biogeocenologii (Fundamentals of forest biogeocenology), Moscow: Nauka, 1964, 573 p.

Targul’yan V. O., Teoriya pedogeneza i evolyucii pochv (Theory of pedogenesis and soil evolution), Moscow: GEOS, 2019, 295 p.

Tomashunas V. M., Abakumov E. V., Organoprofili pochv poluostrovov Yamal i Gydan (Soil organoprofiles of the Yamal and Gydan peninsulas), Mezhdunarodnaya nauchn. konf. «Ekologiya i biologiya pochv» (International Scientific Conference «Ecology and Biology of Soils»), Rostov-na-Donu, 2014, pp. 364–365.

Trofimov S. S., Taranov S. A., Naplekova N. N., Fatkulin F. A., Gumusoobrazovanie v tekhnogennyh ekosistemah (Humus formation in pollution-induced ecosystems), Novosibirsk: Nauka, 1986, 126 p.

Tyurin I. V., Ponomaryova V. V., Materialy po izucheniyu gumusa lesnyh pochv (Materials to the forest soils humus investigation), Trudy Leningradskoj lesotekhnicheskoj akademii, 1940, Vyp. 56, p. 3–49.

Uzhegova I. A., Mahonina G. I., Nachal’nye processy pochvoobrazovaniya na otvalah Pervoural’skogo mestorozhdeniya zheleznyh rud (Initial processes of soil formation on the dumps of the Pervouralsk iron ore deposit), Pochvovedenie, 1984, No 11, pp. 14−21.

Vaksman Z. A., Gumus: Proiskhozhdenie, himicheskij sostav i znachenie ego v prirode (Humus: Origin, chemical composition and its significance in nature), Moscow: Sel’hozgiz, 1937, 471 p.

Vasenev I. I., Pochvennye sukcessii (Soil successions), Moscow: LKI, 2008, 400 p.

Vasenev I. P., Targul’yan V. O., Vetroval i taezhnoe pochvoobrazovanie (Windfall and taiga soil formation), Moscow: Nauka, 1995, 247 p.

Vernadskij V. I., Biosfera (Biosphere) Leningrad: Nauch. him.-tekhnol. izd-vo, 1926, 147 p.

Vernadskij V. I., Zhivoe veshchestvo i biosfera (Living substance and biosphere), Moscow: Nauka, 1994, 671 p.

Wachendorf C., Milbert G., Broll G., Frank T., Graefe U., Beylich A., A concept for a consolidated humus form description – An updated version of the German humus form systematics, International Journal of Plant Biology, 2023, Vol. 14, No 3, pp. 658–686.

Wilde S. A., Forest soils: origin, properties, relation to vegetation and silvicultural management, New York: Wiley & Sons, 1958, 537 p.

Williams V. R., Pochvovedenie. Izbrannye sochineniya (Soil science. Selected works), Moscow: Yurajt, 2025, 344 p.

Yarilov A. A., Pedologiya kak samostoyatel’naya estestvennonauchnaya disciplina o zemle
(Pedology as an independent natural science discipline about the earth), Yur’ev (Tartu): Izd. Yur’evskogo un-ta. 1904 –1905, Part 1, 480 p.; Part 2, 244 p.

Zanella A., Ponge J.-F., Briones M. J. I., Terrestrial humus systems and forms – Biological activity, space-time dynamics, Applied Soil Ecology, 2018, Vol. 122, Part 1, pp. 103–137. DOI: 10.1016/j.apsoil.2017.07.020

Zanella A., Ponge J.-F., Jabiol B., Van Delft B., De Waal R., Katzensteiner K., … & Hager H., Standardized Morpho-Functional Classification of the Planet’s Humipedons, Soil Systems, 2022, Vol. 6, No 3, Article 59, DOI: 10.3390/soilsystems6030059